Frasnian-Famennian (Upper Devonian) transition in the northern hemisphere (NE Laurussia and NE Siberia) - an overview


Authors: Zhuravlev AV, Sokiran EV

Published in: Bulletin of Geosciences, volume 95, issue 4; pages: 419 - 439; Received 18 March 2020; Accepted in revised form 10 July 2020; Online 30 August 2020

Keywords: Frasnian-Famennian boundary, northern hemisphere, conodonts, brachiopods, facies changes, Laurussia, Siberia,

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There is high similarity in sedimentation and taxonomic composition and stratigraphic succession of conodonts and brachiopods across the Frasnian-Famennian (F-F) transition in low- and mid-latitude realms of NE Laurussia and Siberia. This uniformity suggests the presence of low environmental gradients between 10° S and 40° N of the northern hemisphere corresponding to tropical and subtropical climatic zones. The changes in taxonomic diversity in these marine faunas occurred gradually throughout the Frasnian, and were paired with similar, mostly carbonate facies settings, marked by rare development of the Kellwasser-type organic-rich deposits. Moderated ecological effect of the rapid climatic shifts during the F-F crisis is also noteworthy, especially for the shelly benthos, with an exception of the Lilliput effect.


Abramova, A.N. 1999. Frasnian stage at the Western slope of the South Urals. 55 pp. Institute of Geology, Ufa Scientific Center, Russian Academy of Sciences, Ufa. [in Russian]

Alekseev, A.S, Kononova, L.I. & Nikishin, A.M. 1996. The Devonian and Carboniferous of the Moscow Syneclise (Russian Platform): stratigraphy and sea-level changes. Tectonophysics 268, 149-168. View article

Alekseeva, R.E. 1967. Famenskie brakhiopody Severo-Vostoka SSSR, 14-20. In Ivanovskiy, A.B. & Sokolov, B.S. (eds) Novye dannye po biostratigrafii devona i verkhnego paleozoya Sibiri. Sibirskoe otdelenie, Institut Geologii i Geofiziki, Akademiya nauk SSSR. [in Russian]

Alekseeva, R.E., Sidjachenko, A.I., Baranov, V.V., Afanasieva, G.A., Grunt, T.A., Komarov, V.N., Lazarev, S.S. & Manankov, I.N. 1996. Atlas of Devonian Brachiopods from Northeastern Russia (Eastern Yakutia, Magadan Region). 227 pp. Nauka, Moscow. [in Russian]

Aristov, V.A. 1988. Devonskie konodonty Central?nogo Devonskogo Polâ. 120 pp. Nauka, Moscow. [in Russian]

Aristov, V.A. 1994. Devonian and Lower Carboniferous conodonts of the Eurasia: assemblages, zonal subdivision, correlation of different facies sediments. 192 pp. Russian Academy of Sciences, Geological Institute. Moscow. [in Russian]

Averbuch, O., Tribovillard, N., Devleeschouwer, X. & Riquier, L. 2003. Evidences for an orogenic-induced global cooling at the Frasnian-Famennian boundary (ca 376 Ma BP). In EGS AGU EUG Joint Assembly (ed.) Abstracts from the meeting held in Nice, France, 6-11 April 2003, abstract id. 4072.

Baliński, A. 2002. Frasnian-Famenian brachiopod extinction and recovery in Southern Poland. Acta Palaeontologica Polonica 47(2), 289-305.

Baliński, A. & Racki, G. 1999. Wymieranie i odradzanie się faun ramienionogowych a kryzys na granicy franfamen. Przegląd Geologiczny 47, 373-378.

Batanova, G.P. 1955. Brachiopods of the Devonian of the eastern part of the Tatar ASSR. Trudy Vsesouznogo Neftânogo Naučno-Issledovatel’skogo Geologorazvedočnogo Instituta, Novaâ Seriâ 88, 157-195. [in Russian]

Bezgodova, D.V. 2015. A new species of Theodossia nalivkin (Brachiopoda: Spririferida) from the Frasnian of Novaya Zemlya. Trudy Zoologieskogo instituta RAN 319(4), 465-479. [in Russian]

Bond, D.P.G. & Wignall, P.B. 2008. The role of sea-level change and marine anoxia in the Frasnian-Famennian (Late Devonian) mass extinction. Palaeogeography, Palaeoclimatology, Palaeoecology 263, 107-118. View article

Bond, D.P.G., Zaton, M., Wignall, P.B. & Marynowski, L. 2013. Evidence for shallow water ’Upper Kellwasser’ anoxia in the Frasnian-Famennian reefs of Alberta, Canada. Lethaia 46, 355-368. View article

Boucot, A.J., Chen, X. & Scotese, C.R. 2013. Phanerozoic Paleoclimate: An Atlas of Lithologic Indicators of Climate. 478 pp. SEPM Concepts in Sedimentology and Paleontology (Print-on-Demand Version) 11. View article

Carmichael, S.K., Waters, J.A., Königsof, P., Suttner, T.J. & Kido, E. 2019. Paleogeography and paleoenvironments of the Late Devonian Kellwasser event: A review of its sedimentological and geochemical expression, Global and Planetary Change 183, art. 102984. View article

Carmichael, S.K., Waters, J.A., Suttner, T.S., Kido, E. & DeReuil, A.A. 2014. A new model for the Kellwasser Anoxia Events (Late Devonian): Shallow water anoxia in an open oceanic setting in the Central Asian Orogenic belt. Palaeogeography, Palaeoclimatology, Palaeoecology 399, 394-403. View article

Chen, D.Z. & Tucker, M.E. 2003. The Frasnian - Famennian mass extinction: insights from high-resolution sequence stratigraphy and cyclostratigraphy in South China. Palaeogeography, Palaeoecology, Palaeoclimatology 193, 87-111. View article

Copper, P. 1998. Evaluating the Frasnian-Famennian mass extinction: Comparing brachiopod faunas. Acta Palaeontologica Polonica 43(2), 137-154.

Crasquin, S. & Horne, D.J. 2018. The palaeopsychrosphere in the Devonian. Lethaia 51(4), 547-563. View article

Deulin, Yu.V. 2006. Conodonts and correlation of Upper Devonian petroliferous deposits of the Northern Timan-Pechora Province. 253 pp. GIN RAN, Arkhangelsk. [in Russian]

Dzik, J. 2006. The Famennian “Golden Age” of conodonts and ammonoids in the Polish part of the Variscan sea. Palaeontologica Polonica 63, 1-360.

Ellern, S.S., Ivanov, E.E. & Kurbanov, F.J. 1955. Brachiopods of the Devonian of the Tatar ASSR. Trudy Vsesouznogo Neftânogo Naučno-Issledovatel’skogo Geologorazvedočnogo Instituta, Novaâ Seriâ 88, 107-145. [in Russian]

Ernst, R.E. & Youbi, N. 2017. How Large Igneous Provinces affect global climate, sometimes cause mass extinctions, and represent natural markers in the geological record. Palaeogeography, Palaeoecology, Palaeoclimatology 478, 30-52. View article

Fedorova, T.I. 1955. Brachiopods of the Devonian sediments of the Saratov region. Trudy Vsesouznogo Neftânogo Naučno-Issledovatel’skogo Geologorazvedočnogo Instituta, Novaâ Seriâ 88, 7-47. [in Russian]

Gagiev, M.H. 2009. Stratigraphy of Devonian and Lower Carboniferous of the Prikolymian Uplift (Northeastern Asia). 290 pp. NEISRI FEB RAS, Magadan. [in Russian, English abstract]

Gereke, M. & Schindler, E. 2012. “Time-Specific Facies” and biological crises - The Kellwasser Event interval near the Frasnian/Famennian boundary (Late Devonian). Palaeogeography, Palaeoclimatology, Palaeoecology 367-368, 19-29. View article

Gholamalian, H. 2007. Conodont biostratigraphy of the Frasnian-Famennian boundary in the Esfahan and Tabas areas, Central Iran. Geological Quarterly 51(4), 453-476.

Hallam, A. & Wignall, P.B. 1999. Mass extinctions and sea-level changes. Earth-Science Reviews 48(4), 217-250. View article

Hladil, J. 2002. Geophysical records of dispersed weathering products on the Frasnian carbonate platform and early Famennian ramps in Moravia, Czech Republic: proxies for eustasy and palaeoclimate. Palaeogeography, Palaeoclimatology, Palaeoecology 181(1-3), 213-250. View article

Huang, C., Joachimski, M.M. & Gong, Y. 2018. Did climate changes trigger the Late Devonian Kellwasser Crisis? Evidence from a high-resolution conodont δ18OPO4 record from South China. Earth and Planetary Science Letters 495, 174-184. View article

Izokh, O.P., Izokh, N.G., Ponomarchuk, V.A. & Semenova, D.V. 2009. Carbon and oxygen isotopes in the Frasnian-Famennian section of the Kuznetsk Basin (southern West Siberia). Geologiya i Geofizika (Russian Geology and Geophysics) 50(7), 786-795. View article

Joachimski, M.M., Ostertag-Henning, C., Pancost, R.D., Strauss, H., Freeman, K.H., Littke, R., Sinninghe Damsté, J.S. & Racki, G. 2001. Water column anoxia, enhanced productivity and concomitant changes in δ13C and δ34S across the Frasnian-Famennian boundary (Kowala - Holy Cross Mountains/Poland). Chemical Geology 175, 109-131. View article

Kiselev, A.I., Yarmolyuk, V.V., Egorov, K.N., Chernyshov, R.A. & Nikiforov, A.V. 2006. Middle Paleozoic Basic Magmatism of the Northwestern Vilyui Rift: Composition, Sources, and Geodynamics. Petrology 14(6), 588-608. View article

Kravchińsky, V.A. 2012. Paleozoic large igneous provinces of northern Eurasia: Correlation with mass extinction events. Global and Planetary Change 86-87, 31-36. View article

Krawczyński, W., Piechota, A., Sobstel, M., Sokiran, E. & Filipiak, P. 2004. Faunistyczne i środowiskowe zmiany na granicy fran-famen w profilu geologicznym Kamenki (Centralne Pole Dewońskie, Rosja), p. 38. In XIX Konferencja Paleobiologów i Biostratygrafów PTG, Wrocław 16-18.09. 2004.

Krylova, A.K. 1955. Spiriferids of the Devonian of the Volga-Ural region. Trudy Vsesouznogo Neftânogo Naučno-Issledovatel’skogo Geologorazvedočnogo Instituta, Novaâ Seriâ 88, 297-331. [in Russian]

Krylova, A.K. 1959. The Upper Devonian of Stolb Island in the Lena River mouth. Doklady AN SSSR 124(1), 162-164. [in Russian]

Krylova, A.K. 1962. Stratigrafia i brachiopody devona Sibirskoy platformy [Stratigraphy and brachiopods of the Devonian of Siberian Platform]. 108pp. Tr. VNIGRI, 200. Gostoptekhizdat, Leningrad. [in Russian]

Lash, G.G. 2017. A multiproxy analysis of the Frasnian-Famennian transition in western New York State, U.S.A. Palaeogeography, Palaeoclimatology, Palaeoecology 473, 108-122. View article

Ljaschenko, A.I. 1959. Atlas brahiopod i stratigrafiâ devonskih otloženij centralnyh oblastej Russkoj Platformy. 451pp. Gostoptehizdat, Moskva. [in Russian]

Ma, X.P., Becker, R.T., Li, H., & Sun, Y.Y. 2006. Early and Middle Frasnian brachiopod faunas and turnover on the South China shelf. Acta Palaeontologica Polonica 51(4), 789-812.

Ma, X.P., Gong, Y., Chen, D., Racki, G., Chen, X., & Liao, W.H. 2016. The Late Devonian Frasnian-Famennian event in South China - Patterns and causes of extinctions, sea level changes, and isotope variations. Palaeogeography, Palaeoclimatology, Palaeoecology 448, 224-244. View article

Ma, X.P., Sun, Y.L., Hao, W.C., & Liao, W.H. 2002. Rugose corals and brachiopods across the Frasnian - Famennian boundary in central Hunan, South China. Acta Palaeontologica Polonica 47(2), 373-396.

McGhee, G.R. 2013. When the Invasion of Land Failed. The Legacy of the Devonian Extinctions. 336 pp. Columbia University Press, New York. View article

McLaren, D.J., Norris, A.W. & Mcgregor, D.C. 1962. Illustrations of Canadian fossils, Devonian of western Canada. Geological Survey of Canada, Department of Mines and Technical Surveys, paper 62-4, 35 pp. View article

Mikryukov, M.F. 1955. Devonian brachiopods of Western Bashkiria. Trudy Vsesouznogo Neftânogo Naučno-Issledovatel’skogo Geologorazvedočnogo Instituta, Novaâ Seriâ 88, 203-249. [in Russian]

Modzalevskaya, T.L., Alvarez, F. & Rzhonsnitskaya, M.A. 2013. Evolution, migration and biogeography of the plicathyridine brachiopods with a revision of Devonian faunas from the Kuznetsk Basin, Russia. Memoirs of the Association of Australasian Palaeontologists 44, 27-52.

Mottequin, B., Brice, D., Marion, J.-M. & Simon, E. 2016. Plicathyridine brachiopods (Athyridida) from the Frasnian (Late Devonian) of Western Europe and Middle East. Geobios 49(5), 381-393. View article

Nalivkin, D.V. 1947. Class Brachiopoda, 63-134. In Nalivkin, D.V. & Verber, V.N. (eds) Atlas rukovodâsih form iskopaemyh faun SSSR, Vol. 3, Devonskaâ Sistema. VSEGEI-Gosgeoltekhizdat, Leningrad. [in Russian]

Novozhilova, S.I. 1955. Brachiopods of the Devonian of the Kuibyshev and Chkalov Regions. Trudy Vsesouznogo Neftânogo Naučno-Issledovatel’skogo Geologorazvedočnogo Instituta, Novaâ Seriâ 88, 61-89. [in Russian]

Ovnatanova, N.S. & Kononova, L.I. 2008. Frasnian Conodonts from the Eastern Russian Platform. Paleontological Journal 42(10), 997-1166. View article

Ovnatanova, N.S., Kononova, L.I., Kolesnik, L.S. & Gatovsky, Yu.A. 2017. Upper Devonian conodonts of Northeastern European Russia. Paleontological Journal 51(10), 973-1165. View article

Percival, L.M.E., Davies, J.H.F.L., Schaltegger, U., De Vleeschouwer, D., Da Silva, A.-C. & Föllmi, K.B. 2018. Precisely dating the Frasnian-Famennian boundary: implications for the cause of the Late Devonian mass extinction. Scientific Reports 8(1), art. 9578. View article

Pershina, A.I. 1962. Silurian and Devonian deposits of the Chernyshev Swell. 122 pp. Nauka, Moscow, Leningrad. [in Russian]

Pershina, A.I. & Tsyganko, V.S. 1980. New data on Stratigraphy of Devonian and Silurian deposits of the southeast Pai-Khoi, 3-19. In Molin, V.A. & Tsyganko, V.S. (eds) Phanerozoic of the North of the European part of USSR. Syktyvkar. [in Russian]

Pujol, F., Berner, Z., & Stüben, D. 2006. Palaeoenvironmental changes at the Frasnian/Famennian boundary in key European sections: Chemostratigraphic constraints. Palaeogeography, Palaeoclimatology, Palaeoecology 240, 120-145. View article

Qie, W., Algeo, T.J., Luo, G. & Herrmann, A. 2019. Global events of the Late Paleozoic (Early Devonian to Middle Permian): A review. Palaeogeography, Palaeoclimatology, Palaeoecology 531(A), art. 109259. View article

Racki, G. 1998a. Frasnian-Famennian biotic crisis: Undervalued tectonic control? Palaeogeography, Palaeoclimatology, Palaeoecology 141, 177-198. View article

Racki, G. 1998b. The Frasnian-Famennian brachiopod extinction events: A preliminary review. Acta Palaeontologica Polonica 43(2), 395-411.

Racki, G. & Baliński, A. 1998. Late Frasnian Atrypida (Brachiopoda) from Poland and the Frasnian-Famennian biotic crisis. Acta Palaeontologica Polonica 43(2), 273-304.

Racki, G., Racka, M., Matyja, H. & Devleeschouwer, X. 2002. The Frasnian/Famennian boundary interval in the South Polish-Moravian shelf basins: integrated event-stratigraphical approach. Palaeogeography, Palaeoclimatology, Palaeoecology 181, 251-297. View article

Racki, G., Rakocinski, M., Marynowski, L. & Wignall, P.B. 2018. Mercury enrichments and the Frasnian-Famennian biotic crisis: A volcanic trigger proved? Geology 46(6), 543-546. View article

Reddin, C.J., Kocsis, Á.T. & Kiessling, W. 2019. Climate change and the latitudinal selectivity of ancient marine extinctions. Paleobiology 45(1), 70-84. View article

Ricci, J., Quidelleur, X., Pavlov, V., Orlov, S., Shatsillo, A. & Courtillot, V. 2013. New 40Ar/39Ar and K-Ar ages of the Viluy traps (eastern Siberia): Further evidence for a relationship with the Frasnian-Famennian mass extinction. Palaeogeography, Palaeoclimatology, Palaeoecology 386, 531-540. View article

Riquier, L., Averbuch, O., Tribovillard, N., Albani, A. El, Lazreq, N. & Chakiri, S. 2016. Environmental changes at the Frasnian-Famennian boundary in Central Morocco (Northern Gondwana): integrated rock-magnetic and geochemical studies, 197-217. In Becker, R.T. & Kirchgasser, W.T. (eds), Devonian Events and Correlations. Geological Society, London, Special Publications 278. View article

Rodríguez-Canero, R. & Martín-Algarra, A. 2014. Frasnian-Famennian crisis in the Malaguide Complex (Betic Cordillera, Spain). Terra Nova 26(1), 38-54. View article

Rusetskaya, G.A. & Belenitskaya, G.A. 1990. Siberian Platform, 241-253. In Belenitskaya, G.A. & Zadorozhnaya, N.M. (eds) Riftogennye i sul’fatonosnye formatsyi fanerozoâ SSSR. Nedra, Moscow. [in Russian]

Rzhonsnitskaya, M.A. 1968. Devonian Biostratigraphy of the Outskirts of the Kuznetsk Basin. 1. Stratigraphy. 287 pp. Nedra, Leningrad. [in Russian]

Rzhonsnitskaya, M.A. 1975. Devonian Biostratigraphy of the Outskirts of the Kuznetsk Basin. 2. Brachiopod Descriptions 1, Pentamerida and Atrypida. 232 pp. Nedra, Leningrad. [in Russian]

Rzhonsnitskaya, M.A. 1988. Biostratigraphic scheme of the Devonian of the Russian Platform. Canadian Society of Petroleum Geologists, Memoir 14(3), 691-702.

Rzhonsnitskaya, M.A., Markovskii, B.P., Yudina, Y.A. & Sokiran, E.V. 1998. Late Frasnian Atrypida (Brachiopoda) from the South Urals, South Timan and Kuznetsk Basin (Russia). Acta Palaeontologica Polonica 43(2), 305-344.

Sandberg, C.A. & Dreesen, R. 1984. Late Devonian icriodontid biofacies models and alternative shallow-water conodont zonation. Geological Society of America Special Paper 196, 143-178. View article

Sandberg, C.A., Ziegler, W., Dreesen, R. & Butler, J.L. 1988. Late Frasnian Mass Extinction: Conodont Event Stratigraphy, Global Changes, and possible Causes. Courier Forschungsinstitut Senckenberg 102, 263-307.

Schülke, I. 1998. Conodont community structure around the “Kellwasser mass extinction event” (Frasnian/Famennian boundary interval). Senckenbergiana lethaea 77(1/2), 87-99. View article

Scotese, C.R. 2016. PALEOMAP PaleoAtlas for GPlates and the PaleoDataPlotter program. Geological Society of America Abstracts with Programs 48(5). View article

Shen, Z., Song, J., Servais, T. & Gong, Y. 2018. Late Devonian palaeobiogeography of marine organic-walled phytoplankton, Palaeogeography, Palaeoclimatology, Palaeoecology 531(A), art. 108706. View article

Soboleva, M.A., Sobolev, D.B. & Matveeva, N.A. 2018. Razrez franskogo yarusa na r. Kozhym (zapadnyy sklon Pripolyarnogo Urala) - rezul?taty biostratigraficheskikh, bio-, litofatsial?nykh i izotopno-geokhimicheskikh issledovaniy [Frasnian section on the Kozhym River (the western slope of Polar Urals) - results of biostratigraphic, bio- and lithofacies, isotopic and geochemical studies]. Neftegazovaya Geologiya. Teoriya i Praktika 13(1), available at: [in Russian, English abstract] View article

Sokiran, E.V. 2002. Frasnian-Famennian extinction and recovery of rhynchonellid brachiopods from the East European Platform. Acta Palaeontologica Polonica 47(2), 339-354.

Sokiran, E.V. 2006. Early-Middle Frasnian cyrtospiriferid brachiopods from the East European Platform. Acta Palaeontologica Polonica 51(4), 759-772.

Sokolskaja, A.N. 1948. Evolutsia roda Productella i smezhnykh s nim form v paleozoe Podmoskovnoy kotloviny. 168 pp. Nauka, Moscow. [in Russian]

Spalletta, C., Perri, M.C., Over, D.J. & Corradini, C. 2017. Famennian (Upper Devonian) conodont zonation: revised global standard. Bulletin of Geosciences 92(1), 31-57. View article

Stanley, S.M. 2016. Estimates of the magnitudes of major marine mass extinctions in earth history. PNAS 113(42), E6325-E6334. View article

Stigall, A.L. 2010. Invasive species and biodiversity crises: Testing the link in the Late Devonian. PLoS ONE 5(12), e15584. View article

Stigall, A.L. 2012. Speciation collapse and invasive species dynamics during the Late Devonian “Mass Extinction”. GSA Today: A Publication of the Geological Society of America 22(1), 4-10. View article

Stigall, A.L. 2013. Analysing links between biogeography, niche stability and speciation: the impact of complex feedbacks on macroevolutionary patterns. Palaeontology 56(6), 1225-1238. View article

Tsyganko, V.S. 2011. Devon zapadnogo sklona severa Urala i Pai-Khoia (stratigrafiâ, printsipy raschleneniâ, korrelyatsiâ). 356 pp. UrO RAN, Ekaterinburg. [in Russian]

Verneuil, E. 1845. Geologie de la Russie d’Europe et des Montagnes de l’Oural. Paleontologie 2. 512 pp. Paris.

Wang, C. & Wang, Z. 1978. Upper Devonian and Lower Carboniferous conodonts from southern Guizhou. Memoirs of the Nanjing Institute of Geology and Palaeontology, Academia Sinica 11, 51-91.

Winter, J. 2015. Vulkanismus und Kellwasser-Krise-Zirkon-Tephrostratigrafie, Identifizierung und Herkunft distaler Fallout-Aschenlagen (Oberdevon, Synklinorium von Dinant, Rheinisches Schiefergebirge, Harz). Zeitschrift der Deutschen Gesellschaft für Geowissenschaften 166, 227-251. View article

Yazikov, A.Yu., Izokh, N.G., Saraev, S.V., Bakharev, N.K., Gonta, T.V. & Sobolev, E.S. 2013. New data on the Upper Devonian biostratigraphy and sedimentology of Stolb Island (Lena River delta). Russian Geology and Geophysics 54(8), 780-791. View article

Yolkin, E.A., Izokh, N.G. & Romanov, A.P. 2006. New findings of the Early Famennian conodonts from the West Taimyr. News of Paleontology and Stratigraphy 9, 31-37.

Youngquist, W. & Peterson, R.F. 1947. Conodonts from the Sheffield Formation of north-central Iowa. Journal of Paleontology 21, 242-253.

Yudina, Y.A. 1997. Rhynchonellids (Brachiopoda) from the Barma deposits of the western slope of the South Urals. Paleontological Journal 5, 63-67. [in Russian]

Yudina, A.B., Racki, G., Savage, N.S., Racka, M., & Małkowski, K. 2002. The Frasnian-Famennian events in a deep-shelf succession, subpolar Urals: Biotic, depositional and geochemical records. Acta Palaeontologica Polonica 47, 355-372.

Zatoń, M., Zhuravlev, A.V., Rakociński, M., Filipiak, P., Borszcz, T., Krawczyński, W., Wilson, M.A. & Sokiran, E.V. 2014. Microconchid-dominated cobbles from the Upper Devonian of Russia: Opportunism and dominance in a restricted environment following the Frasnian-Famennian biotic crisis. Palaeogeography, Palaeoclimatology, Palaeoecology 401, 142-153. View article

Zhuravlev, A.V. 2008. Stratigraphic characteristic of the Upper Devonian deposits in the Kara River middle course (SW Pay-Khoy). Neftegazovaya Geologiya. Teoriya i Praktika 3(4), available at: [in Russian, English abstract]

Zhuravlev, A.V. 2016. Stroenie i usloviya formirovaniya franskikh otlozheniy severo-zapadnogo Pai-Khoia [Structure and formation conditions of Frasnian section, northwest Pai-Khoi]. Neftegazovaya Geologiya. Teoriya i Praktika 11(4), available at: [in Russian, English abstract] View article

Zhuravlev, A.V., Gerasimova, A.I. & Vevel, Y.A. 2015. Mestnaya stratigraficheskaya schema verkhnedevonsko-srednekamennougol’nogo interval severa Paikhoiskogo karbonatnogo paravtokhtona [Local stratigraphical map of Upper Devonian - Middle Carboniferous interval of Northern Pay Khoy carbonate parautochthone]. Vestnik IG Komi SC UrB RAS 4, 8-13. [in Russian, English abstract] View article

Zhuravlev, A.V. & Smoleva, I.V. 2018. Carbon isotope values in conodont elements from the latest Devonian Early Carboniferous carbonate platform facies (Timan-Pechora Basin). Estonian Journal of Earth Sciences 67(4), 238-246. View article

Ziegler, W. & Lane, H.R. 1987. Cycles in conodont evolution from Devonian to mid-Carboniferous, 147-163. In Aldridge, R.J. (ed.) Palaeobiology of conodonts. Ellis Horwood, Chichester.

Ziegler, W. & Sandberg, C.A. 1990. The Late Devonian standard conodont zonation. Courier Forschungsinstitut Senckenberg 121, 1-115.