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Palaeoecology of a volcanically influenced Silurian sea floor in the Central Prague Basin, and its regional setting
Published in: Bulletin of Geosciences, volume 100, issue 4; pages: 785 - 808; Received 5 November 2025; Accepted in revised form 24 December 2025; Online 31 December 2025
Keywords: palaeoecology, small coral-stromatoporoid build-ups, volcano-sedimentary facies, Ludlow,
Abstract
The palaeoecology of a volcanically influenced Silurian sea floor is analysed from the Smoking Quarry, Kozolupy, Czech Republic. Platy limestones with small coral-stromatoporoid reefs developed around the Svatý Jan volcanic centre in the Prague Basin, alternating with atrypid brachiopod-rich tuffitic shales. The reef building organisms are preserved in situ or very little disturbed, as are the brachiopod assemblages, indicating limited reworking. Extrinsic climatic factors and diagenesis are considered with regard to the sedimentary cyclicity. Limestone firmgrounds and hardgrounds indicate early shallow cementation and exposure of irregular, probably nodular beds enhanced by diagenetic segregation of carbonate. In the volcanic setting, limestones may represent periods of greater current activity, when volcaniclastic sediment was washed into deeper areas, while shales represent more quiescent periods. The Kozolupy section studied represents the uppermost Gorstian volcano-sedimentary facies of the Kopanina Formation going up into shallow water limestones in the early Ludfordian. The succession is compared in its regional context with other sections in the Amerika Anticline, illustrating the spatial development of biofacies along the axis of the Prague Basin and the effects of global eustatic oscillations.References
Barrande, J. 1846. Notice préliminaire sur le Systeme Silurien et les trilobites de Boheme. 97 pp. J.B. Hirschfeld, Leipzig.
Barrande, J. 1865. Systeme silurien du Centre de la Bohéme: Classe de Mollusques, Ordre des Céphalopodes, vol. 2, pt. 1. pls 1-107. Privately published, Prague & Paris.
Barrande, J. 1848. Über die Brachiopoden der silurische Schichten von Boehmen. Naturwlssenschaftliche Abhandlung. W. von Haidinger. Wien. Vol. 2, 357-474, 9 pls.
Barrande, J. 1881 Systeme silurien du centre de la Boheme: Classe des mollusques. Ordre des acéphalés, Vol. 6. 342 pp., 361 pls. C. Bellmann, Prague.
Bek, J., Štorch, P., Tonarová, P. & Libertín, M. 2022. Early Silurian (mid-Sheinwoodian) palynomorphs from the Loděnice-Špičatý vrch, Prague Basin, Czech Republic. Bulletin of Geosciences 97, 385-396.
Bohm, F., Westphal, H. & Bornholdt, S. 2003. Required but disguised: Environmental signals in limestone-marl alternations. Palaeogeography, Palaeoclimatology, Palaeoecology 189, 161-178.
Bouček, B. 1931. On several new fossil rich localities in the Lower Paleozoic of Central Bohemia. Věda přírodní 12.
Boucot, A.J. 1975. Evolution and Extinction Rate Controls. 429 pp. Elsevier, Amsterdam.
Boucot, A.J. & Lawson, J.D. (eds) 1999. Paleocommunities - A Case Study from the Silurian and Lower Devonian. 895 pp. Cambridge University Press, Cambridge, New York, Melbourne.
Buch, L. von 1834. Über Terebratein, mit einem Versuch sie zu classificiren und zu beschreiben. Physikalische Abhandlungen der Königlichen Akademie der Wissenschaften zu Berlin (für 1833) 3, 1-124.
Cherns, L. 1999. Silurian chitons as indicators of rocky shores and lowstand on Gotland, Sweden. Palaios 14, 172-179.
Cherns, L. & Wright, V.P. 2000. Missing molluscs as evidence of large-scale, early skeletal aragonite dissolution in a Silurian sea. Geology 28, 791-794.
Ferretti, A. & Kříž, J. 1995. Cephalopod limestone biofacies in the Silurian of the Prague Basin, Bohemia. Palaios 10, 240-253.
Fiala, F. 1982. Basaltoid diabase from Hostim with indications of hematite mineralisation. Časopis pro mineralogii a geologii 27, 173-185.
Franke, W., Cocks, L.R.M. & Torsvik, T.H. 2017. The Palaeozoic Variscan oceans revisited. Gondwana Research 48, 257-284.
Frýda, J. & Frýdová, B. 2014. First evidence for the Homerian (late Wenlock, Silurian) positive carbon isotope excursion from peri-Gondwana: new data from the Barrandian (Perunica). Bulletin of Geosciences 89, 617-634.
Frýda, J. & Manda, Š. 2013. A long-lasting steady period of isotopically heavy carbon in the late Silurian ocean: evolution of the δ13C record and its significance for an integrated δ13C, graptolite and conodont stratigraphy. Bulletin of Geosciences 88, 463-482.
Frýda, J, Lehnert, O., Frýdová, B., Farkaš, J. &. Kubajko, M. 2021a. Carbon and sulfur cycling during the mid-Ludfordian anomaly and the linkage with the late Silurian Lau/Kozlowskii Bioevent. Palaeogeography Palaeoclimatology Palaeoecology 564, 110152.
Frýda, J., Lehnert, O., Joachimski, M.M., Männik, P., Kubajko, M., Mergl, M., Farkaš, J. & Frýdová, B. 2021b. The Mid-Ludfordian (late Silurian) Glaciation: a link with global changes in ocean chemistry and ecosystem overturns. Earth Science Reviews 220, 103652.
Galle, A. 1973. Family Heliolitidae from the Bohemian Paleozoic. Sborník geologických Věd, Paleontologie 15, 7-48.
Hallam, A. 1986. Origin of minor limestone-shale cycles: climatically induced or diagenetic? Geology 14, 609-612.
Haq, B.U. & Schutter, S.R. 2008. A chronology of Paleozoic sealevel changes. Science 322, 64-68.
Havlíček, V. 1977. Brachiopods of the order Orthida in Czechoslovakia. Rozpravy Ústředního ústavu geologického 44, 1-327.
Havlíček, V. 1984. Lissatrypa (Brachiopoda; Silurian, Lower Devonian) in Bohemia. Věstník ústředního ústavu geologického 59, 217-225.
Havlíček, V. 1995. New data on the distribution of brachiopods in the Motol and lowest Kopanina Formations (Wenlock, lower Ludlow, Prague Basin, Bohemia). Věstník Českého geologického ústavu 70, 47-63.
Havlíček, V. & Štorch, P. 1990. Silurian brachiopods and benthic communities in the Prague Basin (Czechoslovakia), Rozpravy Ústředního ústavu geologického 48, 1-275.
Havlíček, V. & Štorch, P. 1999. Silurian and Lochkovian Communities of the Prague Basin (Barrandian area, Czechoslovakia), 200-228. In Boucot, A.J. & Lawson, J.D. (eds) Paleocommunities - A Case Study from the Silurian and Lower Devonian. Cambridge University Press, Cambridge, New York, Melbourne.
Havlíček, V., Vaněk, J. & Fatka, O. 1994. Perunica microcontinent in the Ordovician (its position within the Mediterranean Province, series division, benthic and pelagic associations). Sborník geologických věd, Geologie 46, 23-56.
Hawle, I. & Corda, A.J.C. 1847. Prodrom einer Monographie der böhmischen Trilobiten. Abhandlungen der königlichen böhmischen Gesellschaft der Wissenschften, V., Folge 5, 119-292.
Horný, R. 1955. The Budňany Beds in the Western Part of the Silurian of the Barrandian. Sborník Ústředního ústavu geologického, Oddíl geologický 21, 315-447.
Horný, R. 1965. Tektonic structure and development of the Silurian between Beroun and Tachlovice. Časopis pro minearalogii a geologii 10, 147-155.
Ivanova, E.A. 1962. Ecology and development of Silurian and Devonian brachiopods of the Kutznetsk, Minussinsk and Tuvinsk basins. Trudy Geologičeskogo instituta, Instituta Akademii nauk SSSR 88, 1-152. [in Russian]
Johnson, M.E. 1996. Stable cratonic sequences and a standard for Silurian eustasy, 203-211. In Witzke, B.J., Ludvigson, G.A. & Day, J. (eds) Paleozoic Sequence Stratigraphy: View from the North American Craton. Geological Society of America Special Paper 306.
Johnson, M.E. 2006. Relationship of Silurian sea-level fluctuations to oceanic episodes and events. GFF 128, 115-121.
Johnson, M.E. 2010. Tracking Silurian eustasy: alignment of empirical evidence or pursuit of deductive reasoning? Palaeogeography, Palaeoclimatology, Palaeoecology 296, 276-284.
Kříž, J. 1984. Autecology and ecogeny of Silurian Bivalvia. Special Papers in Paleontology 32, 183-195.
Kříž, J. 1991. The Silurian of the Prague Basin (Bohemia) - tectonic, eustatic and volcanic controls on facies and faunal development, 179-203. In Bassett, M.G., Lane, P.D. & Edwards, D. (eds) The Murchison Symposium: proceedings of an international conference on The Silurian System. Special Papers in Palaeontology 44.
Kříž, J. 1992. Silurian field excursions: Prague Basin (Barrandian), Bohemia. National Museum of Wales, Geological Series 13, 1-111.
Kříž, J. 1994. Conservation of geological sites, fossils and rock environments in Czechoslovakia. Mémoires de la Société géologique de France 165, 101-102.
Kříž, J. 1998. Recurrent Silurian-lowest Devonian cephalopod limestones of Gondwanan Europe and Perunica, 183-198. In Landing, E. & Johnson, M.E. (eds) Silurian cycles; linkages of dynamic stratigraphy with atmospheric, oceanic, and tectonic changes. Bulletin of New York State Museum 491.
Kříž, J. 1999a. Bivalvia dominated communities of Bohemian type from the Silurian and Lower Devonian carbonate facies, 229-252. In Boucot, A.J. & Lawson, J.D. (eds) Paleocommunities - A Case Study from the Silurian and Lower Devonian. 895 pp. Cambridge University Press, Cambridge, New York, Melbourne.
Kříž, J. 1999b. Silurian Bivalvia - evolution, palaeontology, palaeobiography, importance for biostratigraphy and correlation. Abhandlungen der Geologischen Bundesanstalt 54, 377-384.
Kříž, J. 2008. A new bivalve community from the lower Ludlow of the Prague Basin (Perunica, Bohemia). Bulletin of Geosciences 83, 237-280.
Kříž, J. & Pek, I. 1975. New trace fossils from the Silurian of Bohemia. Časopis pro minearalogii a geologii 20, 175-178.
Kříž, J., Dufka, P., Jaeger, H. & Schönlaub, H.P. 1993. The Wenlock/Ludlow boundary in the Prague Basin (Bohemia). Jahrbuch der Geologische Bundesanstalt 136, 809-839.
Kříž, J., Degardin, J.-M., Ferretti, A., Hansch, W., Gutiérrez Marco, J.C., Paris, F., Almeida, J.M., Robardet, M., Schönlaub, H-P. & Serpagli, E. 2003. Silurian stratigraphy and paleogeography of north Gondwanan and Perunican Europe, 105-178. In Landing, E. & Johnson, M.E. (eds) Silurian lands and seas - paleogeography outside of Laurentia. New York State Museum Bulletin 493.
Krs, M., Krsová, M., Pruner, P., Chvojka, R. & Havlíc˘ek, V. 1987. Palaeomagnetism, palaeogeography and the multicomponent analysis of Middle and Upper Cambrian rocks of the Barrandian in the Bohemian Massif. Tectonophysics 139, 1-20.
Kukal, Z. 1955. Investigation of the Silurian dolomite limestone at Velká Mořina. Sborník Ústředního ústavu geologického, Oddíl geologický 21, 225-251.
Lawson, J.D. 1999. Watkins’ benthic associations in the Ludlow Series of the Welsh Borderland. 388-394. In Boucot, A.J. & Lawson, J.D. (eds) Paleocommunities - A Case Study from the Silurian and Lower Devonian. 895 pp. Cambridge University Press, Cambridge, New York, Melbourne.
Lehnert, O., Frýda, J., Buggisch, W., Munnecke, A., Nützel, A., Kříž, J. & Manda, Š. 2007. δ13C records across the late Silurian Lau event: New data from middle palaeo-latitudes of northern peri-Gondwana (Prague Basin, Czech Republic). Palaeogeography, Palaeoclimatology, Palaeoecology 245, 227-244.
Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. Editio Decima 1. 824 pp. Typis Ioannis Thomae von Trattner, Vindobonae.
Manda, Š. & Kříž, J. 2006. Environmental and biotic changes of the subtropical isolated carbonate platforms during Kozlowskii and Lau events (Prague Basin, Silurian, Ludlow). GFF 128, 161-168.
Melchin, M.J., Sadler, P.M. & Cramer, B.D. 2020. Chapter 21 - The Silurian Period, 695-732. In Gradstein, F.M, Ogg, J.G., Schmitz, M.D. & Ogg, G.M. (eds) Geologic Time Scale 2020. Elsevier.
Michalík, J. 1970. Genus Meristina Hall, 1867 in the Silurian of the Barrandian in Central Bohemia. Sborník geologických věd, Paleontologie 12, 135-170.
Mikuláš, R. 2019. Field trip guidebook, 65-75. In Kočová Veselská, M., Adamovič, J., Kernhoff, M., Rifl, M., Šamánek, J. & Mikuláš, R. (eds) 15th International Ichnofabric Workshop, Prague, Czechia, April 27th - May 3rd, 2019, Program, Abstracts Field Guidebook. The Czech Academy of Sciences, Institute of Geology, Praha.
Milne-Edwards, H. & Haime, J. 1851. Monographie des polypiers fossiles des terrains paléozoiques. Archives du Museum National d’Histoire Naturelle 5, 1-502.
Munnecke, A. & Samtleben, C. 1996. The formation of micritic limestones and the development of limestone-marl alternations in the Silurian of Gotland, Sweden. Facies 34, 159-176.
Murchison R.I. 1839. The Silurian System. 768 pp. John Murray, London.
Nield, E.W. 1982. The earliest Gotland reefs - two bioherms from the Lower Visby Beds (Upper Llandovery). Palaeogeography, Palaeoclimatology, Palaeoecology 39, 149-164.
Nikiforova, O.I. 1970. Brachiopods of the ridge horizon of Vaigach (Late Silurian), 97-144. In Stratigraphy and Fauna of the Silurian Deposits of Vaigach. Naučno-issledovatel’skij institut geologii Arktiki Ministerstva geologii SSSR, Leningrad. [in Russian]
Patočka, F., Pruner, P. & Štorch, P. 2003. Palaeomagnetism and geochemistry of Early Palaeozoic rocks of the Barrandian (Teplá-Barrandian Unit, Bohemian Massif): palaeotectonic implications. Physics and Chemistry of the Earth 28, 735-749.
Rey, Á. del, Frýda, J., Calner, M., Frýdová, B., Zhang, F., Wang, C., Planavsky, N. & Dahl, T.W. 2023. Mid-Ludfordian uranium isotope records distinguish the role of expansive marine anoxia in global carbon cycle dynamics during the late Silurian Lau/Kozlowskii bioevent. Global and Planetary Change 229, 104248.
Riding, R. 1981. Composition, structure and environmental setting of Silurian bioherms in northern Europe, 41-83. In Tommey, D.F. (ed.) European Fossil Reef Models. SEPM Special Publication 30.
Riding, R. 2002. Structure and composition of organic reefs and carbonate mud mounds; concepts and categories. Earth Science Reviews 58, 163-231.
Slavíčková, J. & Kocourek, R. 2000. Clustered trilobites Leonaspis leonhardi in the Kopanina Fomation (Silurian, Czech Republic). Věstník Českého geologického ústavu 75, 153-157.
Šnajdr, M. 1960. Studie o čeledi Scutelluidae (Trilobitae). Rozpravy Ústředního ústavu geologického 26, 1-263.
Šnajdr, M. 1980. Bohemian Silurian and Devonian Proetidae (Trilobita). Rozpravy Ústředního ústavu geologického 45, 1-324.
Sowerby. J.C. de 1818. The mineral conchology of Great Britain 2. B. 251 pp. Meredith, London.
Stel, J.H. 1978. Environment and quantitative morphology of some Silurian tabulates from Gotland. Scripta Geologica 47, 1-75.
Štorch, P. 2023. Graptolite biostratigraphy and biodiversity dynamics in the Silurian System of the Prague Synform (Barrandian area, Czech Republic). Bulletin of Geosciences 98, 1-78.
Štorch, P., Manda, Š. & Loydell, D.K. 2014. The early Ludfordian leintwardinensis graptolite Event and the Gorstian-Ludfordian boundary in Bohemia (Silurian, Czech Republic). Palaeontology 57, 1003-1043.
Tasáryová, Z., Schnabl, P., Čížková, K., Pruner, P., Janoušek, V., Rapprich, V., Štorch, P., Manda, Š., Frýda, J. & Trubač, J. 2014. Gorstian palaeoposition and geotectonic setting of Suchomasty Volcanic Centre (Silurian, Prague Basin, Teplá-Barrandian Unit, Bohemian Massif). GFF 136, 262-265.
Tasáryová, Z., Janoušek, V. & Frýda, J. 2018. Failed Silurian continental rifting at the NW margin of Gondwana: evidence from basaltic volcanism of the Prague Basin (Teplá-Barrandian Unit, Bohemian Massif). International Journal of Earth Sciences 107, 231-1266.
Taylor, A.M. & Goldring, R. 1993. Description and analysis of bioturbation and ichnofabric. Journal of the Geological Society 150, 141-148.
Torsvik, T.H. & Cocks, L.R.M. 2019. The integration of palaeomagnetism, the geological record and mantle tomography in the location of ancient continents. Geological Magazine 156, 242-260.
Vaněk, J. & Vokáč, V. 2004. Trilobitové společenstvo korálových poloh (kopaninské souvrství, ludlow, silur) v “Kouřícím” lomu u Mořiny (Pražská pánev, Barrandien, Čechy). Palaeontologia Bohemiae 9, 1-8.
Watkins, R. 1979. Benthic community organization in the Ludlow Series of the Welsh Borderland. Bulletin of the British Museum Natural History (Geology) 31, 175-280.
Watkins, R. 1996. Paleoecology of Silurian reef bivalves, Racine Formation, North America. Lethaia 29, 171-180.
Weedon, G.P. 1986. Hemipelagic shelf sedimentation and climatic cycles - the basal Jurassic (Blue Lias) Of South Britain. Earth and Planetary Science Letters 76, 321-335.
Weedon, G.P. & Jenkyns, H.C. 1999. Cyclostratigraphy and the Early Jurassic timescale: Data from the Belemnite Marls, Dorset, southern England. Geological Society of America Bulletin 111, 1823-1840.
Westphal, H., Head, M.J. & Munnecke, A. 2000. Differential diagenesis of rhythmic limestone alternations supported by palynological evidence. Journal of Sedimentary Research 70, 715-725.
Westphal, H., Bohm, F. & Bornholdt, S. 2004. Orbital frequencies in the carbonate sedimentary record: distorted by diagenesis? Facies 50, 3-11.
Wright, V.P. & Cherns, L. 2016. How far did feedback between biodiversity and early diagenesis affect the nature of Early Palaeozoic sea floors? Palaeontology 59, 753-765.
Wright, V.P., Cherns L. & Hodges, P. 2003. Missing molluscs: Field testing taphonomic loss in the Mesozoic through early large-scale aragonite dissolution. Geology 31, 211-214.
Wright, V.P., Nohl, T. & Munnecke, A. 2025. Bedding and bedding surfaces in carbonate and mixed carbonate-siliciclastic successions. Part 2, processes, identification and implications of diagenetic bedding, 155-175. In Davies, N.S. & Shillito, A.P. (eds) Bedding Surfaces: True Substrates and Earth’s Historical Archive. Geological Society London, Special Publication 556.
Žák, J. & Sláma, J. 2018. How far did the Cadomian ?terranes’ travel from Gondwana during early Palaeozoic? A critical reappraisal based on detrital zircon geochronology. International Geology Review 60, 319-338.
Žák, J., Kraft, P., Hajná, J., Vacek, F., Svojtka, M., Kachlík, V., Ackerman, L., Verner, K., Zulauf, G., Dörr, W. & Tomek, F. 2025. The Teplá-Barrandian Unit: A fascinating archive recording > 600 My of evolution of European crust, 477-525. In Linneman, U. (ed.) The Variscan Orogen of Central Europe: Geodynamics-Geochronology-Geobiology. Springer, Cham.
