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Early Miocene small mammals from MWQ1/2001 Turtle Joint (Mokrá-Quarry, South Moravia, Czech Republic): biostratigraphical and palaeoecological considerations
Published in: Bulletin of Geosciences, volume 96, issue 1; pages: 99 - 122; Received 28 Mai 2020; Accepted in revised form 25 November 2020; Online 20 December 2020
Keywords: Lagomorpha, Rodentia, Chiroptera, Eulipotyphla, Carpathian Foredeep Basin, Moravian Karst,
Abstract
Mokrá-Quarry (South Moravia, Czech Republic) represents a unique fossil site with an extraordinary abundance and diversity of vertebrate fossil remains. Most research on Mokrá-Quarry localities was focused on herpetofauna. Despite its relevance, the mammal fauna has not been yet studied in detail. In this work, the small mammals from one of the karstic fissures, Mokra´-Western Quarry (1/2001 Turtle Joint), are thoroughly described for the first time, including eight different taxa: Prolagus schnaitheimensis, Prolagus cf. vasconinensis, Rhinolophus cf. cluzeli, Rhinolophus cf. grivensis, Galerix sp., Aliveria aff. luteyni, Megacricetodon sp., and Melissiodon dominans. The Megacricetodon finds represent one of the first appearances of this cricetid in Central Europe and sheds light on the early evolution of the genus. Moreover, the small mammal assemblage confirms an early Miocene age (Burdigalian, MN4) for MWQ1/2001. The paleoenvironment inferred shows a dry karst landscape, with patches of woodlands and open steppe, together with marshy areas.References
Abdul Aziz, H., Böhme, M., Rocholl, A., Prieto, J., Wijbrans, J.R., Bachtadse, V. & Ulbig, A. 2010. Integrated stratigraphy and 40Ar/39Ar chronology of the early to middle Miocene Upper Freshwater Molasse in western Bavaria (Germany). International Journal of Earth Sciences 99, 1859-1886.
Agustí, J. & Antón, M. 2002. Mammoths, Sabertooths, and Hominids. 313 pp. Columbia University Press, New York.
Alba, D.M., Casanovas-Vilar, I., Furió, M., García-Paredes, I., Angelone, C., Jovells-Vaqué, S., Luján, A.H., Almécija, S. & Moya-Sola, S. 2018. Can Pallars i Llobateres: A new hominoid-bearing locality from the late Miocene of the Valles-Penedes Basin (NE Iberian Peninsula). Journal of Human Evolution 121, 193-203.
Álvarez-Sierra, M.A., Daams, R., Lacomba, J.I., Lopez-Martinez, N., van der Meulen, A.J., Sese, C. & de Visser, J. 1991. Palaeontology and biostratigraphy (micromammals) of the continental Oligocene-Miocene deposits of the North-Central Ebro Basin (Huesca, Spain). Scripta Geologica 94, 1-77.
Angelone, C. & Sesé, C. 2009. New characters for species discrimination within the genus Prolagus (Ochotonidae, Lagomorpha, Mammalia). Journal of Paleontology 83(1), 80-88.
Blumenbach, J.F. 1779. Handbuch der Naturgeschichte. 448 pp. J.C.D., Göttingen.
Böhme, M. & Vasylan, D. 2014. Ectothermic vertebrates from the late Middle Miocene of Gratkorn (Austria, Styria). Palaeodiversity and Palaeonvironments 94, 21-40.
Boon-Kristkoiz, E. & Kristkoiz, A.R. 1999. Order Lagomorpha, 259-262. In Rössner, G. & Heissig, K. (eds) The Miocene Land Mammals of Europe. Verlag Dr. Friedrich Pfeil, Munich.
Bosma, A.A., Bruijn, H. de & Wessels, W. 2019. Early and middle Miocene Sciuridae (Mammalia, Rodentia) from Anatolia, Turkey. Journal of Vertebrate Paleontology 38(6), 1-25.
Bowdich, T.E. 1821. An Analysis of the Natural Classifications of Mammalia for the use of students and travelers. 115 pp. J. Smith, Paris.
Brandt, J.F. 1855. Beiträge zur nähern Kenntniss der Säugethiere Russlands. Me´moires de l’Acade´mie impe´riale des sciences de St. Pe´tersbourg 7, 1-365.
Bruijn, H. de 1999. Superfamily Sciuroidea, 271-280. In Rössner, G.E. & Heissig, K. (eds) The Miocene Land Mammals of Europe. Verlag Dr. Friedrich Pfeil, Munich.
Bruijn, H. de, van der Meulen, A.J. & Katsikatsos, G. 1980. The Mammals from the Lower Miocene of Aliveri (lsland of Evia, Greece) Part 1. The Sciuridae. Proceedings B 83(3), 241-261.
Brzobohatý, R. 1997. Paleobatymetrie spodního badenu karpatské předhlubně na Moravě z pohledu otolitovy´ch faun, 37-45. In Hladilová, Š. (ed.) Dynamika vztahů marinního a kontinentálního prostředí. Masaryk Univeristy, Brno.
Brzobohatý, R., Kudělásek, V. & Nehyba, S. 2000. Nejspodnější baden (střední miocén) v okolí Mokré u Brna. Geologické výzkumy na Moravě a ve Slezsku 7, 58-60.
Bulot, C. 1980. Nouvell description de deux especes du genre Megacricetodon (Cricetidae, Rodentia) du Miocene de Bézian (one de La Romieu). Bulletin du Muséum national d’histoire naturelle, Section C Sciences de la terre, paléontologie, géologie, minéralogie 2(1), 3-16.
Cicha, I., Fahlbusch, V. & Fejfar, O. 1972. Die biostratigraphische Korrelation einiger jungtertiärer Wirbeltierfaunen Mitteleuropas. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 140(2), 129-145.
Crusafont, M., Villalta, J.F. De & Truyols, J. 1955. El Burdigaliense continental de la cuenca del Vallés-Penedés. Memorias y Comunicaciones del Instituto Geológico 12, 1-247.
Cuenca-Bescós, G. 1988. Revisión de los Sciuridae del Aragoniense y del Rambliense en la fosa de Calatayud-Montalbán. Scripta Geologica 87, 1-116.
Dehm, R. 1950. Die Nagetiere aus dem Mittel-Miocän (Burdigalium) von Wintershof-West bei Eichstätt in Bayern. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 91, 321-428.
Depéret, C. 1892. Sur la faune d’Oiseaux pliocenes du Roussillon. Mémoires de la Société géologique de France. Paleontologie 3, 127-139.
Díaz Aráez, J.L., Delfino, M., Luján, A.H., Fortuny, J., Bernardini, F. & Alba, D.M. 2017. New remains of Diplocynodon (Crocodylia: Diplocynodontidae) from the early Miocene of the Iberian Peninsula. Comptes Rendus Palevol 16(1), 12-26.
Doukas, C.S. 1986. The mammals from the Lower Miocene of Aliveri (Island of Evia, Greece). Part 5. The insectivores. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen B 89(1), 15-38.
Doukas, C.S. 2003. The MN4 faunas of Aliveri and Karydia (Greece), 127-132. In López-Martínez, N., Peláez-Campomanes, P. & Hernández Fernández, M. (eds) En torno a fósiles de mamíferos: datación, evolución y paleoambiente. Coloquios de Paleontología, Volumen Extraordinario 1.
Engesser, B. 1980. Insectivora und Chiroptera (Mammalia) aus dem Neogen der Türkei. Schweizerische Paläontologische Abhandlungen 102, 49-150.
Ekrt, B., Mikula´š, R., Wagner, J., Cˇerma´k, S., Procha´zkova´, K., Kadlecova´, E. & Fejfar, O. 2016. New contribution to the palaeoichnology and taphonomy of the Ahni´kov fossil site, Early Miocene, Most Basin (the Czech Republic). Fossil Imprint 72(3-4), 202-214.
Estes, R. 1981. Gymmophiona. Caudata, 1-115. In Wellnhofer, P. (ed.) Handbuch der Paläoherpetologie. Munich.
Fahlbusch, V. 1964. Die Cricetiden (Mamm.) der Oberen Su¨ßwasser-Molasse Bayerns. Bayerische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Abhandlungen, Neue Folge 118, 1-136.
Fejfar, O. 1974. Die Eomyiden und Cricetiden (Rodentia, Mammalia) des Miozäns der Tschechoslowakei. Palaeontographica Abteilung A 146, 100-180.
Fejfar, O. 1990. The Neogen VP sites of Czechoslovakia: A contribution to the Neogen Terrestric Biostratigrapphy of Europe Based on Rodents, 211-236. In Lindsay, E.H., Fahlbusch, V. & Mein, P. (eds) European Neogene Mammal Chronology. Plenum Press, New York.
Fejfar, O. & Horáček, I. 1983. Zur Entwicklung der Kleinsäugerfaunen im Villanyium und Alt-Biharium auf dem Gebiet der ČSSR. Schriftenreihe für Geologische Wissenschaften 19-20, 111-208.
Fejfar, O. & Kvaček, Z. 1993. Excursion Nr. 3, Tertiary basins in North-west Bohemia. 35 pp. Universita Karlova and Česká geologická společnost, Prague.
Fejfar, O. & Roček, Z. 1986. The Lower Miocene Vertebrate Fauna of Dolnice, Cheb Basin (Western Bohemia, Czechoslovakia). Acta Universitatis Carolinae, Geologica 2, 233-249.
Fejfar, O., Dvořa´k, Z. & Kadlecova´, E. 2003. New record of Early Miocene (MN3a) mammals in the open brown coal pit Merkur, North Bohemia, Czech Republic, 163-182. In Reumer, J.W.F. & Wessels, W. (eds) Distribution and Migration of Tertiary mammals in Eurasia. A Volume in Honour of Hans de Bruijn. Deinsea 10.
Fischer [de Waldheim], G. 1817. Adversaria zoologica. Me´moires de la Socie´te´ impe´riale des naturalistes de Moscou 5, 357-472.
Freudenthal, M. 1968. On the mammalian fauna of the Hipparion-beds in the Calatayud-Teruel Basin. Part IV: The genus Megacricetodon (Rodentia). Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen B 71, 57-72.
Furió M., Casanovas-Vilar, I. & van den Hoek Ostende, L.W. 2011. Predictable structure of Miocene insectivore (Lipotyphla) faunas in Western Europe along a latitudinal gradient. Palaeogeography, Palaeoclimatology, Palaeoecology 304(3-4), 219-229.
Furió, M., van den Hoek Ostende, L.W., Agustí, J. & Minwer-Barakat, R. 2018. Evolución de las asociaciones de insectívoros (Eulipotyphla, Mammalia) en Espana y su relación con los cambios climáticos del Neógeno y el Cuaternario. Ecosistemas 27(1), 38-51.
Gaillard, C. 1899. Mammiferes miocenes nouveaux ou peu connus de la Grive-Saint-Alban (Isere). Archives du Muséum d’Histoire Naturelle de Lyon 7, 1-78.
García-Alix, A., Minwer-Barakat, R., Martín Suárez, E., Freudenthal, M. & Martín, J. M. 2008. Late Miocene-Early Pliocene climatic evolution of the Granada Basin (southern Spain) deduced from the paleoecology of the micromammal associations. Palaeogeography, Palaeoclimatology, Palaeoecology 265(3-4), 214-225.
Gray, J.E. 1825. An outline of an attempt at the disposition of Mammalia into tribes and families, with a list of the genera apparently appertaining to each tribe. Annals of Philosophy 10, 337-344.
Gregory, W.K. 1910. The orders of mammals. Bulletin of the American Museum of Natural History 27, 1-524.
Gureev, A.A. 1960. Lagomorphs (Lagomorpha) from the Oligocene of Mongolia and Kazakhstan. Trudy Instituta Paleontologii Akademija Nauk SSSR 77, 5-34. [in Russian]
Hand, S. 1985. New Miocene Megadermatids (Chiroptera, Megadermatidae) from Australia with comments on Megadermatid Phylogenetics. Australian Mammalogy 8(1-2), 5-44.
Hilgen, F.J., Lourens, L.J. & Van Dam, J.A. 2012. The Neogene period, 923-978. In Gradstein, F.M., Ogg, J.G., Schmitz, M. & Ogg, G. (eds) The Geologic Time Scale 2012. Elsevier, Amsterdam.
Hladil, J., Bernardová, E., Brunnerová, Z., Brbzobohatý, R., Čekan, V., Dvořák, J., Eliáš, M., Friáková, O., Havlíček, P., Kalvoda, J., Klečák, J., Machatková, B., Maštera, L., Mittrenga, P., Otava, J., Přichystal, A., Rejl, L. & Růžička, M. 1987. Vysvětlivky k základní geologické mapě České republiky 1:25 000 list Mokrá-Horákov. Archive of the Czech Geological Survey. Prague.
Horáček, I. & Špoutil, F. 2012. Why tribosphenic? On variation and constraints in developmental dynamics of chiropteran molars, 410-455. In Gunnell, G.F. & Simmons, N.B. (eds) Evolutionary History of Bats. Fossils, Molecules and Morphology. Cambridge University Press, Cambridge.
Hordijk, K. & van der Meulen, A.J. 2010a. Systematics of resident species of Lagopsis and Prolagus (Ochotonidae, Lagomorpha, Mammalia) from the late early and middle Miocene of northeastern Central Spain, 17-85. In Hordijk, K. Perseverance of pikas in the Miocene. Interplay of climate and competition in the evolution of Spanish Ochotonidae (Lagopmorpha, Mammalia). Ph.D. thesis, Utrecht University.
Hordijk, K. & van der Meulen, A.J. 2010b. Systematics of immigrant species of Prolagus (Ochotonidae, Lagomorpha, Mammalia) from the late early and middle Miocene of north-eastern Central Spain, 87-141. In Hordijk, K. Perseverance of pikas in the Miocene. Interplay of climate and competition in the evolution of Spanish Ochotonidae (Lagopmorpha, Mammalia). Ph.D. thesis, Utrecht University.
Hordijk, K., Duijnstee, I.A.P. & van der Meulen, A.J. 2010. Speciation, evolution and competition in pikas (Ochotonidae, Lagomorpha, Mammalia) from the early and middle Miocene of Southwestern Europe, 143-173. In Hordijk, K. Perseverance of pikas in the Miocene. Interplay of climate and competition in the evolution of Spanish Ochotonidae (Lagopmorpha, Mammalia). Ph.D. thesis, Utrecht University.
Hordijk, K., Bosma, A., Bruijn, H. de, van Dam, J., Geraedts, C., van den Hoek Ostende, L.W., Reumer, J. & Wessels, W. 2015. Biostratigraphical and palaeoecological implications of the small mammal assemblage from the late early Miocene of Montalvos 2, Teruel Basin, Spain. Palaeobiodiversity and Palaeoenvironments 95(3), 321-346.
Hugueney, M. 1965. Les Chiropteres du Stampien supérieur de Coderet-Branssat (Allier). Travaux et Documents des Laboratoires de Géologie de Lyon 9(1), 97-127.
Ivanov, M. 2008. Early Miocene Amphibians (Caudata, Salientia) from the Mokrá-Western Quarry (Czech Republic) with comments on the evolution of Early Miocene amphibian assemblages in Central Europe. Geobios 41(4), 465-492.
Ivanov, M. & Musil, R. 2004. Prˇedbeˇžné výsledky výzkumu neogenních obratlovcu° z lokality Mokrá-lom. Acta Musei Moraviae Scientiae Geologicae 89, 223-236.
Ivanov, M., Musil, R. & Brzobohatý, R. 2006. Terrestrial and Marine Faunas from the Miocene Deposits of the Mokra´ Plateau (Drahany Upland, Czech Republic) - Impact on Palaeogeography. Beiträge zur Paläontologie 30, 223-239.
Ivanov, M., Ruta, M., Klembara, J. & Böhme, M. 2018. A new species of Varanus (Anguimorpha: Varanidae) from the early Miocene of the Czech Republic, and its relationships and palaeoecology. Journal of Systematic Palaeontology 16(9), 767-797.
Ivanov, M., Čerňanský, A., Bonilla-Salomón, I. & Luján, A.H. 2020. Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications. Geodiversitas 42(20), 343-376.
Jovells-Vaqué, S. & Casanovas-Vilar, I. 2018. A review of the genus Melissiodon (Cricetidae, Rodentia) in the Valles-Penede`s Basin (Catalonia). Journal of Vertebrate Paleontology 38(5), 1-9.
Kälin, D. 1999. Tribe Cricetini, 373-387. In Rössner, G.E. & Heissig, K. (eds) The Miocene Land Mammals of Europe. Verlag Dr. Friedrich Pfeil, Munich.
Kälin, D. & Kempf, O. 2009. High-resolution stratigraphy from the continental record of the Middle Miocene Northern Alpine Foreland Basin of Switzerland. Neues Jahrbuch Fur Geologie Und Palaontologie, Abhandlungen 254(1-2), 177-235.
König, C.D.E. 1825. Icones. Fossilium Sectiles: Centuria Prima. 4 pp. G.B. Sowerby, London.
Lacépede, B.G.E. 1799. Mémoire sur une nouvelle table méthodique des animaux a mamelles. Mémoires de l’Institut National des Sciences et des Arts, Sciences Mathématiques et Physiques 3, 469-502.
Lartet, E. 1851. Notice sur la colline de Sansan, Suivie d’une Récapitulation de Diverses Especes d’Animaux Vertébrés Fossiles Trouvés soit a Sansan, soit dans d’Autres Gisements du Terrain Tertiaire Miocene dans le Bassin Sous-Pyrénéen. 45 pp. J.-A. Portes, Auch.
Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum Classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus 1. Edito decima, reformata. 824pp. Laurentii Salvii, Holmiae.
López-Martínez, N. 1989. Revisión sistemática y biostratigráfica de los Lagomorpha (Mammalia) del Terciario y Cuaternario de Espana. Memorias del Museo Paleontologico de la Universidad de Zaragoza 3, 1-342.
López-Martínez, N. 1997. Lignées évolutives, formes intermédiaires et évolution en mosaique: exemple de lagomophes européens. In Aguilar J.P., Legendre, S. & Michaux, J. (eds) Actes du Congres BiochroM’97. Mémoires et Travaux de l’Institut de Montpellier de l’Ecole Pratique Hautes Etudes 21, 105-120.
López-Martínez, N. 2001. Paleobiogeographical history of Prolagus, an European ochotonid (Lagomorpha). Lynx 32, 215-231.
Luján, A.H., Delfino, M., Casanovas-Vilar, I. & Alba, D.M. 2014. Taxonomy of subgenus Temnoclemmys Bergounioux, 1958 (Testudines: Geoemydidae: Ptychogasterinae) based on new material from the Valles-Penedes Basin. Comptes Rendus Paleovol 13(4), 277-295.
Luján, A.H., Delfino, M., Robles, J.M. & Alba, D.M. 2016. A review of the Miocene tortoise Testudo catalaunica Bataller, 1926 and a revised phylogeny of extinct species of genus Testudo (Testudines: Testudinidae). Zoological Journal of the Linnean Society 178(2), 312-342.
Luján, A.H., Ivanov, M., Fortuny, J. & Sabol, M. 2017. Fossil turtle remains from the early Miocene (MN4) localities of Mokra´ Quarry (Czech Republic), 86. In YNHM: Young Natural History scientists Meeting. Paris.
Luján, A.H., Chroust, M., Čerňanský, A., Fortuny, J., Mazuch, M. & Ivanov, M. 2019. First record of Diplocynodon ratelii Pomel, 1847 from the early Miocene site of Tušimice (Most Basin, Northwest Bohemia, Czech Republic). Comptes Rendus Palevol 18(7), 877-889.
Luján, A.H., Čerňanský, A., Bonilla-Salomón, I., Březina, J. & Ivanov, M. in press. Fossil turtles from the early Miocene localities of the Mokrá-Quarry (Burdigalian, MN4), Moravia, Czech Republic. Geodiversitas.
Markovic´, Z., Bruijn, H. de & Wessels, W. 2016. A revision of the new rodent collections from the Early Miocene of Sibnica, Serbia, 63-117. In Markovic´, Z. & Milivojevic´, M. (eds) Life on the Shore - Geological and Paleontological Research in the Neogene of Sibnica and Vicinity (Levacˇ basin, Central Serbia). Part 1. Special Issue of the Natural History Museum in Belgrade. Natural History Museum, Belgrade.
Matthew, W.D. & Granger, W. 1923. Nine new rodents from the Oligocene of Mongolia. American Museum Novitates 102, 1-10.
Mein, P. 1958. Les mammiferes de la faune sidérolithique de Vieux-Collonges. Nouvelles Archives du Muséum d’Histoire Naturelle de Lyon 5(1), 3-122.
Mein, P. 1989. Die Kleinsa¨ugerfauna des Untermioza¨ns (Eggenburgien) von Maigen, Niedero¨sterreich. Annalen des Naturhistorischen Museums in Wien 90 (A), 49-58.
Mein, P. & Freudenthal, M. 1971. Une nouvelle classification des Cricetidae (Rodentia, Mammalia) du Tertiaire de l’Europe. Scripta Geologica 2, 1-37.
Mein, P. & Freudenthal, M. 1981. Les Cricetidae (Mammalia, Rodentia) du Néogene Moyen de Vieux-Collonges. Scripta Geologica 60, 1-11.
Minwer-Barakat, R., García-Alix, A., Agustí, J., Martín Suárez, E. & Freudenthal, M. 2009. The micromammal fauna from Negratín-1 (Guadix Basin, Southern Spain): new evidence of African-Iberian mammal exchanges during the Late Miocene. Journal of Paleontology 83(6), 854-879.
Oliver, A. & Peláez-Campomanes, P. 2013. Megacricetodon vandermeuleni, sp. nov. (Rodentia, Mammalia), from the Spanish Miocene: a new evolutionary framework for Megacricetodon. Journal of Vertebrate Paleontology 33(4), 943-955.
Oliver, A. & Peláez-Campomanes, P. 2014. Evolutionary patterns of early and middle Aragonian (Miocene) of Megacricetodon (Rodentia, Mammalia) from Spain. Palaeontographica Abteilung A 303, 85-135.
Oliver, A. & Peláez-Campomanes, P. 2016. Early Miocene evolution of the genus Megacricetodon in Europe and its paleobiogeographical implications. Acta Palaeontologica Polonica 61(1), 211-219.
Peláez-Campomanes, P. & Daams, R. 2002. Middle Miocene rodents from Paşalar, Anatolia, Turkey. Acta Palaeontologica Polonica 47(1), 125-132.
Pomel, A. 1848. Etudés sur les carnassiers insectivores (extrait). Seconde partie, Classification des insectivores. Archives des sciences physiques et naturelles 9, 244-251.
Pomel, A. 1853. Catalogue méthodique et descriptif des vertébrés fossiles: découverts dans le bassin hydrographique supérieur de la Loire, et surtout dans la vallee de son affluent principal, l’Allier. 193 pp. J.-B. Bailliere, Paris.
Procházka, V.J. 1899. Miocénové ostrovy v krasu Moravském. Rozpravy České akademie 2, 37.
Qiu, Z. 1996. Middle Miocene Micromammals Faunas from Tunggur, Nei, Mongolia. 216 pp. Beijing Science Press, Beijing.
Qiu, Z., Li, C. & Wang, S. 1981. Miocene mammalian fossils from Xining Basin, Qinghai. Vertebrata Palasiatica 19(2), 156-173.
Reichenbacher, B., Krijgsman, W., Lataster, Y., Pipperr, M., Van Baak, C.G.C., Chang, L., Kälin, D., Jost, J., Doppler, G., Jung, D., Prieto, J., Abdul Aziz, H., Böhme, M., Garnish, J., Kirscher, U. & Bachtadse, V. 2013. A new magnetostratigraphic framework for the Lower Miocene (Burdigalian/Ottnangian, Karpatian) in the North Alpine Foreland Basin. Swiss Journal of Geosciences 106, 309-334.
Revilliod, P. 1920. Contribution a l’étude des chiropteres des terrains tertiaries: 2eme partie. Mémoires de la Société Paléontologique Suisse 44, 63-129.
Rez, J. 2003. Strukturní analy´za jižní části Moravského krasu v lomech Mokrá a okolí. 45 pp. Master thesis, Masaryk University, Brno.
Rez, J. 2010. Strukturně-geologický vývoj jižní části Moravského krasu. 75 pp. Ph.D. thesis, Masaryk University, Brno.
Ringeade, M. 1979. Decouverte de nouveaux Lagomorphes dans le Miocene inferieur d’Aquitaine et implications biostratigraphiques. Bulletin de l’Institut de Geologie du Bassin d’Aquitaine 26, 111-157.
Roman, F. & Viret, J. 1930. Le Miocene continental de l’Armagnac et le gisement Burdigalien de La Romieu (Gers). Livre Jubilaire de la Société Géologique de France 2, 577-604.
Rössner, G.E. 2004. Community structure and regional patterns in late Early to Middle Miocene Ruminantia of Central Europe. In Steininger, F.F., Kovar-Eder, J. & Fortelius, M. (eds) The Middle Miocene Environments and Ecosystem Dynamics of the Eurasian Neogene (EEDEN). Courier Forschungsinstitut Senckenberg 249, 91-100.
Sabol, M., Fejfar, O., Horáček, I. & Čermák, S. 2007. The Early Miocene micromammalian assemblage from Mokra´ - 1/2001 Turtle Joint site (Moravia, Czech Republic) - preliminary results. Scripta Facultatis Scientiarum Naturalium Universitatis Masarykianae Brunensis 36, 57-64.
Schaub, S. 1920. Melissiodon n. gen., ein bisher u¨bersehener oligoca¨ner Muride. Senckenbergiana 2, 43-47.
Schaub, S. 1925. Die Hamsterartigen Nagetiere des Tertia¨rs und ihre lebenden Verwandten. Abhandlungen der Schweizerischen Pala¨ontologischen Gesellschaft 45, 1-110.
Steininger, F.F. 1999. Chronostratigraphy, Geochronology and Biochronology of the Miocene “European Land Mammal Mega-Zones” (ELMMZ) and the Miocene “Mammal-Zones (MN-Zones)”, 9-24. In Rössner, G.E. & Heissig, K. (eds) The Miocene Land Mammals of Europe. Verlag Dr. Friedrich Pfeil, Munich.
Storch, G. 1999. Order Chiroptera, 81-90. In Rössner G.E. & Heissig, K. (eds) The Miocene Land Mammals of Europe. Verlag Dr. Friedrich Pfeil, Munich.
Tobien, H. 1975. Zur Gebissstruktur, Systematik und Evolution der Genera Piezodus, Prolagus und Ptychoprolagus (Lagomorpha, Mammalia) aus einingen Vorkommen im jüngeren Tertiär Mittel- und Westeuropas. Notizblatt des Hessischen Landesamtes für Bodenforschung zu Wiesbaden 103, 103-186.
van den Hoek Ostende, L.W. & Fejfar, O. 2006. Erinaceidae and Talpidae (Erinaceomorpha, Soricomorpha, Mammalia) from the Lower Miocene of Merkur-Nord (Czech Republic, MN3). Beiträge zur Paläontologie 30, 175-203.
van der Meulen, A.J., García-Paredes, I., Álvarez-Sierra, M.Á., Van den Hoek Ostende, L.W., Hordijk, K., Oliver, A., López-Guerrero, P., Hernández-Ballarín, V. & Peláez-Campomanes, P. 2011. Biostratigraphy or biochronology? Lessons from the Early and Middle Miocene small Mammal Events in Europe. Geobios 44(2-3), 309-321.
Vasileiadou, K. & Koufos, G.D. 2005. The micromammals from the Early/Middle Miocene locality of Antonios, Chalkidiki, Greece. Annales de Paléontologie 91(3), 197-225.
Venczel, M. 2004. Middle Miocene anurans from the Carpathian Basin. Palaeontographica Abteilung A 271(5-6), 151-174.
Villa, A., Abella, J., Alba, D.M., Almécija, S., Bolet, A., Koufos, G.D., Knoll, F., Luján, A.H., Morales, J., Robles, J.M., Sánchez, I.M. & Delfino, M. 2018. Revision of Varanus marathonensis (Squamata, Varanidae) based on historical and new material: morphology, systematics, and paleobiogeography of the European monitor lizards. PLoS ONE 13, e0207719.
Wessels, W., Weerd, A.A., Bruijn, H. de & Marković, Z. 2018. New Melissiodontinae (Mammalia, Rodentia) from the Paleogene of south-east Serbia. Palaeobiodiversity and Palaeoenvironments 98(3), 1-17.
Zapfe, H. 1950. Die Fauna der miozänen Spaltenfüllung von Neudorf an der March (ČSR.). Chiroptera. Sitzungsberichte Österreichischen der Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, Abt. I 159, 51-64.
Ziegler, R. 1990. Didelphidae, Erinaceidae, Metacodontidae und Dimylidae (Mammalia) aus dem Oberoligozän und Untermiozän Süddeutschlands. Stuttgarter Beiträge zur Naturkunde 158 (B), 1-99.
Ziegler, R. 1993. Die Chiroptera (Mammalia) aus dem Untermiozän von Wintershof- West bei Eichstätt (Bayern). Mitteilungen der Bayerischen Staatssammlung für Paläontologie und Historische Geologie 33, 119-154.
Ziegler, R. 1999. Order Insectivora, 53-74. In Rössner, G.E. & Heissig, K. (eds) The Miocene Land Mammals of Europe. Verlag Dr. Friedrich Pfeil, Munich.
Ziegler, R. 2003. Bats (Chiroptera, Mammalia) from Middle Miocene karstic fissure fillings of Petersbuch near Eichstätt, Southern Franconian Alb (Bavaria). Geobios 36(4), 447-490.
Ziegler, R. & Fahlbusch, V. 1986. Kleinsäuger-Faunen aus der basalen Oberen Süßwasser-Molasse Niederbayerns. Zitteliana 14, 3-58.