A synopsis of Westphalian?earliest Stephanian medullosalean and allied plant fossils from the Central and Western Bohemian basins, Czech Republic


Authors: Šimůnek Z, Cleal CJ

Published in: Bulletin of Geosciences, volume 95, issue 4; pages: 441 - 468; Received 22 January 2020; Accepted in revised form 5 June 2020; Online 18 October 2020

Keywords: Medullosan pteridosperms, Pennsylvanian, Radnice Member, Nýřany Member, Westphalian,

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A revised data of the medullosalean foliage remains from the CWBB



The Medullosales were among the most diverse seed-plant clades in the Pennsylvanian palaeotropical, peat-forming wetlands and their fossil foliage constitutes valuable biostratigraphical and palaeofloristic indices. However, there have been several recent taxonomic revisions of the intramontane floras of the Central and Western Bohemian basins, as well as a number of unpublished records that were not incorporated into previous palaeogeographical studies. In this paper, we collate all of the currently available data on medullosalean foliage from the middle and upper Pennsylvanian Kladno Formation from these basins. The different autochthonous and allochthonous floras of the Radnice Member were coded separately to see if the habitats that represent may have had an effect on the analysis. The new dataset suggests that the roof shale floras, although still distinct from those of the lowland paralic basins, are more comparable to the similar aged intramontane Saar-Lorraine floras than was suggested previously. The apparent palaeofloristic difference between the Central and Western Bohemia and Saar-Lorraine floras seems to have been at least partly because the data for the former area included species from both ash deposits that representing peat substrate vegetation and roof shale floras representing clastic substrate vegetation.


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Wagner, R.H. 1962. A brief review of the stratigraphy and floral succession of the Carboniferous in NW. Spain. Compte rendu 4e Congres International de Stratigraphie et de Géologie du Carbonifere (Heerlen) 3, 753-762.

Wagner, R.H. 1963. Sur les Callipteridium du Westphalien supérieur et du Stéphanien. Comptes rendus de l?Académie des Sciences, Paris 257, 719-721.

Wagner, R.H. 1964a. Stephanian floras in NW. Spain, with special reference to the Westphalian D - Stephanian A boundary. Compte rendu 5e Congres International de Stratigraphie et de Géologie du Carbonifere (Paris, 1963) 1, 835-851.

Wagner, R.H. 1964b. Stephanian B flora from the Cinera-Matallana Coalfield (León) and neighbouring outliers. I: Mixoneura, Reticulopteris, Linopteris and Odontopteris. Notas y Communicaciones del Instituto Geológico y Minero de Espana 75, 5-56.

Wagner, R.H. 1966. Palaeobotanical dating of Upper Carboniferous folding phases in NW. Spain. Memorias del Instituto Geologico y Minero de Espana 66, 1-169.

Wagner, R.H. 1968. Upper Westphalian and Stephanian species of Alethopteris from Europe, Asia Minor and North America. Mededelingen van de Rijks Geologische Dienst, Serie C, III-1 6, 1-188.

Wagner, R.H. 1974. The chronostratigraphic units of the Upper Carboniferous in Europe. Bulletin de la Société Belge de Géologie, de Paléontologie et d?Hydrologie 83, 235-253.

Wagner, R.H. 1977. Comments on the upper Westphalian and Stephanian floras of Czechoslovakia, with particular reference to their stratigraphic age, 441-457. In Holub, V.M. & Wagner, R.H (eds) Symposium on Carbononiferous Stratigraphy. Geological Survey, Prague.

Wagner, R.H. 1984. Megafloral zones of the Carboniferous. Compte rendu 9e Congres International de Stratigraphie et de Géologie du Carbonifere (Washington, 1979) 2, 109-134.

Wagner, R.H. & Álvarez-Vázquez, C. 2008. A revision of the Lower Pennsylvanian Alethopteris lonchitica (auctorum) and its identity with Alethopteris urophylla. Revista Espanola de Paleontología 23, 157-192.

Wagner, R.H. & Castro, M.P. 1998. Neuropteris obtusa, a rare but widespread Late Carboniferous pteridosperm. Palaeontology 41, 1-22.

Wagner, R.H., Fernández Garcia, L.G. & Eagar, R.M.C. 1983. Geology and palaeontology of the Guardo Coalfield (NE Leon - NW Palencia), Cantabrian Mts. 109 pp. Ministerio de Industria y Energia, Madrid.

Wittry, J., Glasspool, I.J., Béthoux, O., Koll, R & Cleal, C.J. 2015. A revision of the Pennsylvanian marattialean fern Lobatopteris vestita auct. and related species. Journal of Systematic Palaeontology 13, 615-643. View article

Zeiller, R. 1899. Étude sur la flore fossile du basin houiller d’Héraclée (Asie Mineure). Mémoires de la Société Géologique de France, Paléontologie 21, 1-91. View article

Zodrow, E.L. & Cleal, C.J. 1998. Revision of the pteridosperm foliage Alethopteris and Lonchopteridium (Upper Carboniferous), Sydney Coalfield, Nova Scotia, Canada. Palaeontographica, Abteilung B 247, 65-122.

Zodrow, E.L. & McCandlish, K. 1978. Distribution of Linopteris obliqua in the Sydney Coalfield of Cape Breton Island, Nova Scotia. Palaeontographica, Abteilung B 168, 1-16.

Zodrow, E.L., Tenchov, E.L. & Cleal, C.J. 2007. The arborescent Linopteris obliqua plant (Medullosales, Pennsylvanian). Bulletin of Geosciences 82, 51-84. View article