Ostracods of the Toarcian (Jurassic) of Peniche, Portugal: taxonomy and evolution across and beyond the GSSP interval


Authors: Cabral MC, Lord AR, Pinto S, Duarte LV, Azeredo AC

Published in: Bulletin of Geosciences, volume 95, issue 3; pages: 243 - 278; Received 20 December 2019; Accepted in revised form 27 May 2020; Online 4 July 2020

Keywords: Pliensbachian-Toarcian, ostracods, CIE, taxonomy, biostratigraphy, palaeobiogeography, Portugal,

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The stratigraphical occurrence of ostracod assemblages of the uppermost Pliensbachian (emaciatum Biozone) and Lower-Middle Toarcian (polymorphum, levisoni and bifrons biozones) at Peniche, western Portugal, is reported, an interval that encompasses the Toarcian Global Boundary Stratotype Section and Point (GSSP) and the Toarcian Oceanic Anoxic Event as represented by a Carbon Isotope Excursion (CIE). The broad pattern of ostracod faunal turnover and replacement in the Peniche GSSP section before, during and after the CIE, including “survivor” and “Lazarus” species, and the stepwise extinction of metacopine ostracods is similar to that described from elsewhere in Western Europe. Assemblage composition is closest to material described from the Fastnet Basin, offshore SW Ireland, and the Mochras Farm Borehole, Wales, Great Britain. The δ13C record at Peniche and Mochras Farm facilitates a very refined comparison of ranges of species in common between the two sites. The species Bairdia paramolesta sp. nov. and Kinkelinella ventrocarinata sp. nov. are described.


Adamczak, F. 1967. Morphology of two Silurian metacope ostracodes from Gotland. Geologiska Föreningens i Stockholm Förhandlingar 88, 462-475.View article

Adamczak, F. 1976. Morphology and Carapace Ultrastructure of some Healdiidae (Ostracoda). Abhandlungen und Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg, New Series 18/19 (Suppl.), 315-318.

Ainsworth, N.R. 1986. Toarcian and Aalenian Ostracoda from the Fastnet Basin, offshore southwest Ireland. Geological Survey of Ireland Bulletin 3, 277-336.

Ainsworth, N.R. 1987. Pliensbachian Ostracoda from the Fastnet Basin, offshore Southwest Ireland. Geological Survey of Ireland Bulletin 4, 41-62.

Ainsworth, N.R. 1989. Remarks on the nomenclature of two ostracod species from the Pliensbachian and Upper Toarcian-Aalenian of the Fastnet Basin, offshore southwest Ireland. Geological Survey of Ireland Bulletin 4, 165-166.

Ainsworth, N.R. & Boomer, I. 2009. Lower Jurassic (Hettangian-Toarcian), 175-197. In Whittaker, J.E. & Hart, M.B. (eds) Ostracods in British Stratigraphy. The Micropalaeontological Society Special Publication. Geological Society, London.

Andreu, B., Qajoun, A. & Cubaynes, R. 1995. Ostracodes du Toarcien du Quercy (Bassin d’Aquitaine, France): systématique, biostratigraphie et paléobiogéographie. Geobios 28, 209-240.View article

Apostolescu, V. 1959. Ostracodes du Lias du Bassin de Paris. Revue de l’Institut Français du Petrole 14, 795-826.

Arias, C. 2006. Changes in ostracod provincialism during the early Toarcian in the European epicontinental sea-western Tethys area. Revista Espanola de Micropaleontología 38, 245-267.

Arias, C. 2009. Extinction pattern of marine Ostracoda across the Pliensbachian-Toarcian boundary in the Cordillera Ibérica, NE Spain: Causes and consequences. Geobios 42, 1-15.View article

Arias, C. & Lord, A.R. 1999a. Upper Pliensbachian and Lower Toarcian Ostracoda from the Cordillera Ibérica, north-east Spain. Part I. Revista Espanola de Micropaleontologia 31, 73-98.

Arias, C. & Lord, A.R. 1999b. Upper Pliensbachian and Lower Toarcian Ostracoda from the Cordillera Ibérica, north-east Spain. Part II. Revista Espanola de Micropaleontologia 31, 219-242.

Azeredo, A.C., Duarte, L.V., Henriques, M.H. & Manuppella, G. 2003. Da dinâmica continental no Triásico aos mares no Jurássico Inferior e Médio. 43 pp. Cadernos de Geologia de Portugal, Instituto Geológico e Mineiro, Lisboa.

Baird, W. 1845. Arrangement of the British Entomostraca, with a list of species, particularly noticing those which have as yet been discovered within the bounds of the Club. Transactions of the Berwickshire Naturalists’ Club 2, 145-158.

Baird, W. 1850. The Natural History of the British Entomostraca. 364 pp. The Ray Society, London.View article

Bate, R.H. 1963. Middle Jurassic Ostracoda from North Lincolnshire. Bulletin of the British Museum (Natural History), Geology 8, 173-219.

Bate, R.H. 2009. Middle Jurassic (Aalenian-Bathonian), 175-197. In Whittaker, J.E. & Hart, M.B. (eds) Ostracods in British Stratigraphy. The Micropalaeontological Society Special Publication. Geological Society, London.

Bate, R.H. & Coleman, B.E. 1975. Upper Lias Ostracoda from Rutland and Huntingdonshire. Bulletin of the Geological Survey of Great Britain 55, 1-42.

Bizon, J.-J. 1960. Sur quelques Ostracodes du Lias du Bassin Parisien. Revue de Micropaléontologie 2, 203-211.

Blake, J.F. 1876. Subclass Entomostraca, 429-435. In Tate, R. & Blake, J.F. The Yorkshire Lias. 475 pp. John Van Voorst, London.

Boomer, I. 1988. On Ektyphocythere lanceolata Boomer sp. nov. A Stereo-Atlas of Ostracod Shells 15, 89-92.

Boomer, I. 1991. Lower Jurassic ostracod biozonation of the Mochras Borehole. Journal of Micropalaeontology 9, 205-218.View article

Boomer, I. 1992. Lower Jurassic ostracods from Ilminster, Somerset, England. Journal of Micropalaeontology 11, 47-57.View article

Boomer, I. & Ainsworth, N.R. 2009. Lower Jurassic (Hettangian-Toarcian), 175-197. In Whittaker, J.E. & Hart, M.B. (eds) Ostracods in British Stratigraphy. The Micropalaeontological Society Special Publication. Geological Society, London.

Boomer, I., Ainsworth, N.R. & Exton, J. 1998. A re-examination of the Pliensbachian and Toarcian Ostracoda of Zambujal, west-central Portugal. Journal of Micropalaeontology 17, 1-14.View article

Boomer, I., Lord, A. & Crasquin, S. 2008. The extinction of the Metacopina (Ostracoda). Senckenbergiana lethaea 88, 47-53.View article

Boomer, I., Lord, A.R., Page, K.H., Bown, P.R., Lowry, F.M.D. & Riding, J.B. 2009. The biostratigraphy of the Upper Pliensbachian-Toarcian (Lower Jurassic) sequence at Ilminster, Somerset. Journal of Micropalaeontology 28, 67-86.View article

Boutakiout, M., Donze, P. & Oumalch, F. 1982. Nouvelles especes d’ostracodes du Lias moyen et supérieur du Jbel Dhar En Nsour (Rides sud-Rifaines, Maroc Septentrionale). Revue de Micropaléontologie 25, 94-104.

Brady, G.S. 1867. A synopsis of the Recent British Ostracoda. The Intellectual Observer 12, 110-130.

Bradfield, H.H. 1935. Pennsylvanian Ostracoda of the Ardmore Basin, Oklahoma. Bulletins of American Paleontology 22(73), 1-173.

Brand, E. 1961. In Brand, E. & Malz, H. Ostracoden-studien im Dogger, 3: drei neue Procytheridea-arten und Ljubimovella n.g. aus dem NW-Deutschen Bajocien. Senckenbergiana lethaea 42, 157-173.

Cabral, M.C., Colin, J.-P., Azeredo, A.C., Silva, R.L. & Duarte, L.V. 2015. Brackish and marine ostracode assemblages from the Sinemurian of western Portugal, with description of new species. Micropaleontology 61, 3-24.

Cabral, M.C., Lord, A.R., Boomer, I.D., Loureiro, I. & Malz, H. 2014. Tanycythere new genus and its significance for Jurassic ostracod diversity. Journal of Paleontology 88, 519-530.View article

Cabral, M.C., Loureiro, I., Duarte, L.V. & Azeredo, A.C. 2013. Registo da extinçao dos Metacopina (Ostracoda, Crustacea) no Toarciano de Rabaçal, regiao de Coimbra. Comunicaçoes Geológicas 100, Especial I, 63-68.

Cabral, M.C., Loureiro, I.M., Pinto, S., Duarte, L.V. & Azeredo, A.C. 2011. New light on the Metacopina (Crustacea, Ostracoda) Toarcian extinction event: integrated data from the Lusitanian Basin, Portugal. Geophysical Research Abstracts 13, art. EGU2011-3607.

Caruthers, A.H., Smith, P.L. & Gröcke, D.R. 2013. The Pliensbachian-Toarcian (Early Jurassic) extinction, a global multi-phased event. Palaeogeography, Palaeoclimatology, Palaeoecology 386, 104-118.View article

Caswell, B.A., Coe, A.L. & Cohen, A.S. 2009. New range data for marine invertebrate species across the early Toarcian (Early Jurassic) mass extinction. Journal of the Geological Society of London 166, 859-872.View article

Comas-Rengifo, M.J., Duarte, L.V., Felix, F.F., Joral, F.G., Goy, A. & Rocha, R.B. 2015. Latest Pliensbachian-Early Toarcian brachiopod assemblages from the Peniche section (Portugal) and their correlation. Episodes 38, 2-8.View article

Correia, V.F., Riding, J.B., Duarte, L.V., Fernandes, P. & Pereira, Z. 2017. The palynological response to the Toarcian Oceanic Anoxic Event (Early Jurassic) at Peniche, Lusitanian Basin, western Portugal. Marine Micropaleontolology 137, 46-63.View article

Coryell, H.N. 1963. Bibliographic index and classification of the Mesozoic Ostracoda. 1175 pp. University of Dayton Press, Dayton.

Coryell, H.N. & Fields, S. 1937. A Gatun Ostracode fauna from Cativa, Panama. American Museum Novitates 956, 1-18.

Dana, J.D. 1852. Crustacea: Cyproidea. United States Exploring Expedition during the Years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U.S.N. 13(2), 1277-1304.

Danise, S., Twitchett, R.J., Little, C.T.S. & Clémence, M.E. 2013. Marine benthic community dynamics through the Early Toarcian (Early Jurassic) global warming event. PLoS One 8, art. e56255.View article

Dera, G., Brigaud, B., Monna, F., Laffont, R., Pucéat, E., Deconinck, J.-K., Pellenard, P., Joachimski, M.M. & Durlet, C. 2011. Climatic ups and downs in a disturbed Jurassic world. Geology 39, 39-51.View article

Drexler, E. 1958. Foraminiferen und Ostracoden aus dem Lias α von Siebeldingen/Pfalz. Geologisches Jahrbuch 75, 475-554.

Dreyer, E. 1967. Mikrofossilien des Rät und Lias von SW-Brandenburg. Jahrbuch für Geologie 1(1965), 491-531.

Duarte, L.V. 1997. Facies analysis and sequential evolution of the Toarcian-Lower Aalenian series in the Lusitanian Basin (Portugal). Comunicaçoes do Instituto Geológico e Mineiro 83, 65-94.

Duarte, L.V. 2007. Lithostratigraphy, sequence stratigraphy and depositional setting of the Pliensbachian and Toarcian series in the Lusitanian Basin (Portugal). Ciencias da Terra (UNL) 16, 17-23.

Duarte, L.V. & Soares, A.F. 2002. Litostratigrafia das séries margo-calcárias do Jurássico inferior da Bacia Lusitânica (Portugal). Comunicaçoes do Instituto Geológico e Mineiro 89, 135-154.

Duarte, L.V., Comas-Rengifo, M.J., Hesselbo, S., Mattioli, E., Suan, G., Baker, S., Cabral, M.C., Correia, V., García Joral, F., Goy, A., Reolid, M., Rita, P., Félix, F., Paredes, R., Pittet, B. & Rocha, R.B. 2018. The Toarcian oceanic anoxic event at Peniche. An exercise in integrated stratigraphy - stop 1.3, 33-54. In Duarte, L.V. & Silva, R.L. (eds) 2nd International Workshop on the Toarcian Oceanic Anoxic Event: Field Trip Guidebook, Coimbra, September 7th-8th, 2018-IGCP-655: The Toarcian Oceanic Anoxic Event in the Western Iberian Margin and its Context within the Lower Jurassic Evolution of the Lusitanian Basin. University of Coimbra.

Duarte, L.V., Silva, R.L., Félix, F., Comas-Rengifo, M.J., Rocha, R.B., Mattioli, E., Paredes, R., Mendonça Filho, J.G. & Cabral, M.C. 2017. The Jurassic of the Peniche Peninsula (Portugal): scientific, educational and science popularization relevance. Revista de la Sociedad Geológica de Espana 30, 55-70.

Duarte, L.V., Silva, R.L., Oliveira, L.C.V., Comas-Rengifo, M.J. & Silva, F. 2010. Organic-rich facies in the Sinemurian and Pliensbachian of the Lusitanian Basin, Portugal: Total Organic Carbon distribution and relation to transgressive-regressive facies cycles. Geologica Acta 8, 325-340.

Duarte, L.V., Wright, V.P., López, S.F., Elmi, S., Krautter, M., Azeredo, A.C., Henriques, M.H., Rodrigues, R. & Perilli, N. 2004. Early Jurassic carbonate evolution in the Lusitanian Basin (Portugal): Facies, sequence stratigraphy and cyclicity, 45-71. In Duarte, L.V. & Henriques, M.H. (eds) Carboniferous and Jurassic Carbonate Platforms of Iberia. Universidade de Coimbra. 23rd International Association of Sedimentologists Meeting. Field Trip Guidebook 1.

Exton, J. 1979. Pliensbachian and Toarcian Microfauna of Zambujal, Portugal: Systematic Paleontology. Carleton University Geological Paper 79-1, 1-103.

Exton, J. & Gradstein, F.M. 1984. Early Jurassic stratigraphy and micropalaeontology of the Grand Banks and Portugal, 13-30. In Westermann, G.E.G. (ed.) Jurassic-Cretaceous Biochronology and Paleogeography of North America. Geological Association of Canada Special Paper 27.

Fantasia, A., Adette, T., Spangenberg, J.E., Font, E., Duarte, L.V. & Föllmi, K.B. 2019. Global versus local processes during the Pliensbachian-Toarcian transition at the Peniche GSSP, Portugal: A multi-proxy record. Earth-Science Reviews 198, art. 102932. View article

Field, R.A. 1966. Species of the family Cytherellidae (Ostracoda) from the Lower Lias of South Dorset, England. Senckenbergiana lethaea 47, 87-105.

Fischer, W. 1962. Ostracoden der Gattung Monoceratina Roth 1928, Cytheropteron G.O. Sars 1865 und andere im Lias zeta Schwabens. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 114, 333-345.

Franz, M., Tesakova, E. & Beher, E. 2009. Documentation and revision of the index ostracods from the Lower and Middle Jurassic in SW Germany according to Buck (1954). Palaeodiversity 2, 119-167.

Gómez, J.J. & Arias, C. 2010. Rapid warming and ostracods mass extinction at the Lower Toarcian (Jurassic) of central Spain. Marine Micropaleontology 74, 119-135.View article

Gómez, J.J. & Goy, A. 2011. Warming-driven mass extinction in the Early Toarcian (Early Jurassic) of northern and central Spain. Correlation with other time-equivalent European sections. Palaeogeography, Palaeoclimatology, Palaeoecology 306, 176-195.View article

Gómez, J.J., Goy, A. & Canales, M.L. 2008. Seawater temperature and carbon isotope variations in belemnites linked to mass extinction during the Toarcian (Eartly Jurassic) in Central and Northern Spain. Comparison with other European sections. Palaeogeography, Palaeoclimatology, Palaeoecology 258, 28-58.View article

Gradstein, F.M., Ogg, J.G., Schmitz, M.D. & Ogg, G.M. (eds) 2012. The Geological Timescale 2012. 1176 pp. Elsevier B.V.

Gramann, F. 1963. Liasina n. gen. (Ostracoda) aus dem Deutschen Lias. Geologisches Jahrbuch 82, 65-73.

Gründel, J. 1964. Zur gattung Healdia (Ostracoda) und zu einigen verwandten formen aus dem Unteren Jura. Geologie 13, 456-477.

Gründel, J. 1967. Zur Großgliederung der Ordnung Podocopida G.W. Müller, 1894 (Ostracoda). Neues Jahrbuch für Geologie und Paläontologie, Monatshefte 1967, 321-332.

Gründel, J. & Kozur, H. 1972. Zur taxonomie der Bythocytheridae und Tricornidae (Podocopida, Ostracoda). Monatsberichte der Deutschen Akademie der Wissenschaften zu Berlin 13 (for 1971), 907-937.

Hallam, A. 1961. Cyclothems, Transgressions and Faunal Change in the Lias of North-West Europe. Transactions of the Edinburgh Geological Society 18, 124-174.View article

Harlton, B.H. 1933. Micropaleontology of the Pennsylvanian Johns Valley Shale of the Ouachita Mountains, Oklahoma, and its relationship to the Mississippian Caney Shale. Journal of Paleontology 7, 3-29.

Herrig, E. 1979. Die Gattung Bairdia (Ostracoda, Crustacea) im Lias von Thüringen. Teil I. Zeitschrift für Geologische Wissenschaften 7, 641-661.

Herrig, E. 1985. Zur Taxonomie und Evolution der Gattung Kinkelinella Martin, 1960 (Ostracoda) im Unteren Jura von Mittel- und Nordwesteuropa. Zeitschrift für Geologische Wissenschaften 13, 715-723.

Hesselbo, S.P., Jenkyns. H.C., Duarte, L.V. & Oliveira, L.C.V. 2007. Carbon-isotope record of the Early Jurassic (Toarcian) Oceanic Anoxic Event from fossil wood and marine carbonate (Luistanian Basin, Portugal). Earth and Planetary Science Letters 253, 455-470.View article

Issler, A. 1908. Beiträge zur Stratigraphie und Mikrofauna des Lias in Schweben. Palaeontographica 55, 1-104.

Jenkyns, H.C. 1985. The Early Toarcian and Cenomanian-Turonian anoxic events in Europe: comparisons and contrasts. Geologisches Rundschau 74, 505-518.View article

Jenkyns, H.C. 1988. The early Toarcian (Jurassic) anoxic event: stratigraphic, sedimentary, and geochemical evidence. American Journal of Science 288, 101-151.View article

Jenkyns, H.C. 2010. Geochemistry of oceanic anoxic events. Geochemistry, Geophysics, Geosystems 11, art. Q03004.View article

Jones, T.R. 1849. A monograph of the Entomostraca of the Cretaceous Formation of England. Annual Volumes (Monographs) of the Palaeontographical Society 3, 1-40.View article

Jones, T.R. & Sherborn, C.D. 1888. On some Ostracoda from the Fullers-earth Oolite and Bradford Clay. Proceedings of the Bath Natural History and Field Club 6, 249-278.

Kaufmann, A. 1900. Cypriden und Darwinuliden der Schweiz. Revue Suisse de Zoologie 8, 209-423.View article

Knitter, H. 1983. Biostratigraphische Untersuchungen mit Ostracoden im Toarcien Süddeutschlands. Facies 8, 213-262.View article

Knitter, H. & Riegraf, W. 1984. Biostratigraphie (Cephalopoden, Ostracoden) des Oberen Toarcium von Blumberg-Achdorf/Wutach und Weilheim/Teck (Baden-Württemberg). Jahrbuch des Geologisches Landesamt Baden-Württemberg 26, 57-97.

Kollmann, K. 1960. Ostracoden aus der Alpinen Trias Österreichs - 1: Parabairdia n.g. und Ptychobairdia n.g. (Bairdiidae). Jahrbuch der Geologischen Bundesanstalt, Sonderband 5, 79-105.

Korte, C., Hesselbo, S.P., Ullmann, C.V., Dietl, G., Ruhl, M., Schweigert, G. & Thibault, N. 2015. Jurassic climate mode governed by ocean gateway. Nature Communications 6, art. 10015.View article

Kozur, H. 1972. Die Bedeutung triassischer Ostracoden für stratigraphische und paläoökologische Untersuchungen. Mitteilungen der Gesellschaft der Geologie und Bergbaustudenten in Oesterreich 21, 623-660.

Kristan-Tollmann, E. 1977. Zur Evolution des Schließmuskelfeldes bei Healdiidae und Cytherellidae (Ostracoda). Neues Jahrbuch für Geologie und Paläontologie, Monatshefte 1977, 621-639.

Kristan-Tollmann, E. 1993. Zur paläogeographischen Verbreitung der Ostracoden-Gattung Hermiella an der Rhät/Lias-Grenze. Zitteliana 20, 331-342.

Latreille, P.A. 1806. Genera Crustaceorum et Insectorum secundum ordinem naturalem in familias disposita. 302 pp. Amand Koenig, Paris.View article

Little, C.T.S. & Benton, M.J. 1995. Early Jurassic mass extinctions: a global long-term event. Geology 23, 495-498.View article

Littler, K., Hesselbo, S.P. & Jenkyns, H.C. 2010. A carbon-isotope perturbation at the Pliensbachian-Toarcian boundary: evidence from the Lias Group, NE England. Geological Magazine 147, 181-192.View article

Ljubimova, P.S. 1955. Ostrakody Mezozoyskikh otlozheniy srednego Povolzhya i Obshchego Syrta. Trudy Vsesoyuznogo Neftyango Nauchno-Issledovatelskogo Geologo-razvedochnogo Instituta (VNIGRI), Novaya Seriya 84, 3-189.

Lord, A.R. 1982. Metacopine ostracods in the Lower Jurassic, 262-277. In Banner, F.T. & Lord, A.R. (eds) Aspects of Micropalaeontology. George Allen and Unwin, London.View article

Lord, A. & Moorley, A. 1974. On Ogmoconcha ambo Lord and Moorley sp. nov. A Stereo-Atlas of Ostracod Shells 2, 9-16.

Lord, A.R., Cabral, M.C. & Danielopol, D.L. 2020. Sieve-type normal pore canals in Jurassic ostracods: A review with description of a new genus. Acta Palaeontologica Polonica 65, 37 pp.View article

Malz, H. 1971. Zur Taxonomie ‘glattschaliger’ Lias-Ostracoden. Senckenbergiana lethaea 52, 433-455.

Mandelstam, M.I. 1959. Ostrakody iz otlozheniy Paleogena sredney Azii. Trudy Vsesoyuznogo Neftyango Nauchno-Issledovatelskogo Geologo-razvedochnogo Instituta (VNIGRI), Novaya Seriya 136 (Mikrofauna SSSR: 10), 442-542.

Mandelstam, M.I. 1960. In Orlov, Y.A. (ed.) Osnovy Paleontologii 8, 365-411.

Martin, G.P.R. 1960. In Hoffmann, K. & Martin, G.P.R. Die zone des Dactylioceras tenuicostatum (Toarcien, Lias) in NW- und SW-Deutschland. Paläontologische Zeitschrift 34, 103-149.View article

Mattioli, E., Pittet, B., Petitpierre, L. & Mailliot, S. 2009. Dramatic decrease of pelagic carbonate production by nannoplankton across the Early Toarcian anoxic event (TOAE). Global and Planetary Change 65(3-4), 134-145.View article

M’Coy, F. 1844. A synopsis of the characters of the Carboniferous Limestone fossils of Ireland. 207 pp. University Press, Dublin.View article

McKenzie, K.G. 1968. Saipanetta, new name for Saipanella McKenzie, 1967 (Ostracoda, Podocopida) non Saipanella Chamberlin, 1945 (Myriopoda). Crustaceana 14, 104-105.View article

Monostori, M. 2008. Comparative study of the Pliensbachian and Toarcian ostracods in the Gerecse and Mecsek Mountains, Hungary. Hantkeniana 6, 17-31.

Mouterde, R. 1955. Le Lias de Peniche. Comunicaçoes dos Serviços Geológicos de Portugal 36, 87-115.

Müller, G.W. 1894. Die Ostracoden des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. Fauna und Flora des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. Herausgegeben von der Zoologischen Station zu Neapel 21, 1-404.

Müller, G.W. 1912. Ostracoda in das Tierreich. Das Tierreich 31, 1-434.

Norman, A.M. 1865. Report on the Crustacea of the deep sea dredging off the coasts of Northumberland and Durham. Transactions of the Natural History Society of Northumberland, Durham and Newcastle-upon-Tyne 1, 12-19.

Ogg, J., Ogg, G. & Gradstein, F. 2016. A Concise Geologic Time Scale. 240 pp. Elsevier.

Olempska, E., Horne, D.J. & Szaniawaski, H. 2012. First record of preserved soft parts in a Palaeozoic podocopid (Metacopina) ostracod, Cytherellina submagna: phylogenetic implications. Proceedings of the Royal Society of London B, 279, 564-570.View article

Pais, V., Cabral, M.C., Lord, A., Azeredo, A.C. & Gallagher, L. 2016. Middle Jurassic Tethyan-Boreal ostracod faunal links: a case study from the Callovian of Portugal. Journal of Micropalaeontology 35, 205-228.View article

Pinto, S. 2008. Ostracodos do Toarciano da Bacia Lusitânica (Peniche e Alvaiázere). 202 pp. Master thesis, Universidade de Lisboa, Faculdade de Ciencias, Departamento de Geologia, Portugal.

Pinto, S., Cabral, M.C. & Duarte, L.V. 2007. Preliminary data on the ostracod fauna from the Lower Toarcian of Peniche. Ciencias da Terra (UNL) 16, 37-43.

Pittet, B., Suan, G., Lenoir, F., Duarte, L.V. & Mattioli, E. 2014. Carbon isotope evidence for sedimentary discontinuities in the Lower Toarcian of the Lusitanian Basin (Portugal): sea level change at the onset of the Oceanic Anoxic Event. Sedimentary Geology 303, 1-14.View article

Quenstedt, F.A. 1858. Der Jura. 842 pp. Tübingen, Laupp & Siebeck.

Richter, H. 1987. Die Mikrofauna des Unteren Toarciums der Nordschweiz. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 176, 137-155.

Riegraf, W. 1984. Neue Ostracoden-Arten aus dem Oberen Pliensbachium und Unteren Toarcium Südwestdeutschlands und Südfrankreichs. Stuttgarter Beiträge zur Naturkunde B, 104, 1-19.

Riegraf, W. 1985. Mikrofauna, Biostratigraphie und Fazies im Unteren Toarcium Südwestdeutschlands und Vergleiche mit benachbarten Gebieten. Tübinger Mikropaläontologische Mitteilungen 3, 1-232.

Rita, P., Reolid, M. & Duarte, L.V. 2016. Benthic foraminiferal assemblages record major environmental perturbations during the Late Pliensbachian-Early Toarcian interval in the Peniche GSSP, Portugal. Palaeogeography, Palaeoclimatology, Palaeoecology 454, 267-281.View article

Rocha, R.B., Mattioli, E., Duarte, L.V., Pittet, B., Elmi, S., Mouterde, R., Cabral, M.C., Comas-Rengifo, M.J., Gómez, J.J., Goy, A., Hesselbo, S.P., Jenkyns, H.C., Littler, K., Mailliot, S., Oliveira, L.C.V., Osete, M.L., Perilli, N., Pinto, S., Ruget, C. & Suan, G. 2016. Base of the Toarcian Stage of the Lower Jurassic defined by the Global Boundary Stratotype Section and Point (GSSP) at the Peniche section (Portugal). Episodes 39, 460-481.View article

Rodríguez-Tovar, F.J., Miguez-Salas, O. & Duarte, L.V. 2017. Toarcian Oceanic Anoxic Event induced unusual behaviour and palaeobiological changes in Thalassinoides tracemaker. Palaeogeography, Palaeoclimatology, Palaeoecology 485, 46-56.View article

Roemer, F.A. 1841. Die versteinerungen des Norddeutschen Kreidegebirges. 145 pp. Hannover.View article

Roth, R. 1928. Monoceratina: a new genus of Ostracoda from the Pennsylvanian of Oklahoma. Journal of Paleontology 2, 15-19.

Ruebsam, W., Mayer, B. & Schwark, L. 2019. Cryosphere carbon dynamics control early Toarcian global warming and sea level evolution. Global and Planetary Change 172, 440-453.View article

Sars, G.O. 1866. Oversigt af Norges marine ostracoder. Forhandlinger i Videnskabs-selskebet i Christiania 1865, 1-130.

Sars, G.O. 1887. Nye bidrag til kundskaben om Middelhavets invertebratfauna: 4. Ostracoda Mediterranea (Sydeuropaeiske). Archiv for Mathematik og Naturvidenskab 12, 173-324.View article

Sars, G.O. 1926. An account of the Crustacea of Norway with short descriptions and figures of all the species. Volume 9, parts 13-14. 209-240 pp. Bergen Museum, Bergen.

Schmid-Röhl, A., Röhl, H.J., Oschmann, W., Frimmel, A. & Schwark, L. 2002. Palaeoenvironmental reconstruction of Lower Toarcian epicontinental black shales (Posidonia Shale, SW Germany): global versus regional control. Geobios 35, 13-20.View article

Shaver, R.H. 1961. In Benson, R.H. et al. Systematic Descriptions, Q99-Q414. In Moore, R.C. (ed.) Treatise on Invertebrate Paleontology, Q. Arthropoda 3, Crustacea, Ostracoda. Geological Society of America, New York.

Stephenson, M.B. 1946. Weches Eocene Ostracoda from Smithville, Texas. Journal of Paleontology 20, 297-344.

Suan, G., Mattioli, E., Pittet, B., Lécuyer, C., Suchéras-Marx, B., Duarte, L.V., Philippe, M., Reggiani, L. & Martineau, F. 2010. Secular environmental precursors to Early Toarcian (Jurassic) extreme climate changes. Earth and Planetary Science Letters 290, 448-458.View article

Suan, G., Mattioli, E., Pittet, B., Mailliot, S. & Lécuyer, C. 2008. Evidence for major environmental perturbation prior to and during the Toarcian (Early Jurassic) oceanic anoxic event from the Lusitanian Basin, Portugal. Paleoceanography 23, art. PA1202.View article

Sylvester-Bradley, P.C. 1961. In Benson, R.H. et al. Systematic Descriptions, Q99-Q414. In Moore, R.C. (ed.) Treatise on Invertebrate Paleontology, Q. Arthropoda 3, Crustacea, Ostracoda. Geological Society of America, New York.

Tate, R. & Blake, J.F. 1876. The Yorkshire Lias. 475 pp. John Van Voorst, London.

Trecalli, A., Spangenberg, J., Adatte, T., Föllmi, K.B. & Parente, M. 2012. Carbonate platform evidence of ocean acidification at the onset of the early Toarcian oceanic anoxic event. Earth and Planetary Science Letters 357, 214-225.View article

Triebel, E. 1941. Zur Morphologie und Ökologie der fossilen Ostracoden. Mit beschreibung einiger neuer gattungen und arten. Senckenbergiana 23, 294-400.

Triebel, E. 1950. Die taxonomische Stellung dere Ostracoden-Gattung Ogmoconcha und der Lectotypus von O. amalthei. Senckenbergiana 31, 113-120.

Triebel, E. & Bartenstein, H. 1938. Die Ostracoden des deutschen Juras. I. Monoceratina-Arten aus dem Lias und Dogger. Senckenbergiana 20, 502-518.

Van Den Bold, W.A. 1960. Eocene and Oligocene Ostracoda of Trinidad. Micropaleontology 6, 145-196.View article

Wright, V.P. & Wilson, R.C.L. 1984. A carbonate submarine-fan sequence from the Jurassic of Portugal. Journal of Sedimentary Petrology 54, 394-412.View article

Xu, W., Ruhl, M., Jenkyns, H.C., Hesselbo, S.P., Riding, J.B., Selby, D., Naafs, B.D.A., Weijers, J.W.H., Pancost, R.D., Tegelaar, E.W. & Idiz, E.F. 2017. Carbon sequestration in an expanded lake system during the Toarcian Oceanic Anoxic Event. Nature Geoscience 10, 129-134.View article

Xu, W., Ruhl, M., Jenkyns, H.C., Leng, M.J., Huggett, J.M., Minisini, D., Ullmann, C.V., Riding, J.B., Weijers, J.W.H., Storm, M.S., Percival, L.M.E., Tosca, N.J., Idiz, E.F., Tegelaar, E.W. & Hesselbo, S.P. 2018. Evolution of the Toarcian (Early Jurassic) carbon-cycle and global climatic controls on local sedimentary processes (Cardigan Bay Basin, UK). Earth and Planetary Science Letters 484, 396-411.View article