The phacopine trilobite genera Morocops Basse, 2006 and Adrisiops gen. nov. from the Devonian of Morocco

 

Authors: Viersen AP van, Holland D, Koppka J

Published in: Bulletin of Geosciences, volume 92, issue 1; pages: 13 - 30; Received 10 June 2016; Accepted in revised form 25 January 2017; Online 20 February 2017

Keywords: Trilobita, Phacopidae, taxonomy, palaeobiogeography, Morocco, Algeria, Emsian, Eifelian,

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Abstract

Well-preserved specimens of the phacopine trilobites Morocops Basse (= Barrandeops McKellar & Chatterton) and Adrisiops gen. nov. are recorded from upper Emsian strata in Morocco. Morocops is a problematic taxon; the close similarities of its stratigraphically youngest members to the earliest members of Geesops Struve are indicative of grade taxonomy. So far, Morocops is exclusively Gondwanan although potential congeners have been described from peri-Gondwana. One of the stratigraphically oldest species of Geesops occurs in the lower Eifelian of the French Ardennes (Laurussia). Its co-occurrence with pioneer “Bohemian” trilobites there suggests that faunal exchange between Laurussia and peri-Gondwanan terranes was unhindered at that time. New material of Morocops torkozensis (Schraut) is recorded; however, the type locality and horizon of this species remain uncertain. Adrisiops gen. nov. Is erected to accommodate strongly vaulted, Gondwanan taxa that share a unique cephalic morphology inclusive of a short, but rather inflated glabella and ventrally deflected anterior border. New taxa are Morocops spinifer sp. nov. And Adrisiops weugi gen. et sp. nov.

References

ABOUSSALAM, Z.S., BECKER, R.T. & BULTYNCK, P. 2015. Emsian (Lower Devonian) conodont stratigraphy and correlation of the Anti-Atlas (Southern Morocco). Bulletin of Geosciences 90, 893-980.View article

ALBERTI, G.K.B. 1970. Trilobiten des jüngeren Siluriums sowie des Unter- und Mittel-Devons. II. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 525, 1-233.

ALBERTI, G.K.B. 1983. Trilobiten des jüngeren Siluriums sowie des Unter- und Mittel-Devons. IV. Senckenbergiana lethaea 64, 1-88.

ATTEMS, C.M.T. VON 1951. Myriopoden aus Höhlen des Belgischen Congo. Revue de zoologie et de botanique africaines 44, 386-404.

BARRANDE, J. 1852. Systeme Silurien du centre de la Bohęme. Iere partie. Recherches paléontologiques. I. Trilobites. 935 pp. Prague and Paris.View article

BASSE, M. 1998. Trilobiten aus mittlerem Devon des Rhenohercynikums: III. Proetida (3), Phacopida (2), Lichida (Lichoidea, Odontopleuroidea) und ergänzende Daten. Palaeontographica (A) 249, 1-162.

BASSE,M. 2006. Eifel-Trilobiten IV. Proetida (3), Phacopida (3). 304 pp. Quelle & Meyer-Verlag, Wiebelsheim.

BASSE,M. 2012. Fossilium Catalogus I: Animalia. Pars 150. Trilobites Africae: Catalogus typorum. 311 pp. Margraf Publishers GmbH, Weikersheim.

BECKER, R.T., ABOUSSALAM, S.Z., BOCKWINKEL, J., EBBIGHAUSEN, V., EL HASSANI, A. & NÜBEL, H. 2004a. Upper Emsian stratigraphy at Rich Tamelougou near Torkoz (SW Dra Valley, Morocco). Documents de l’Institut Scientifique, Rabat 19, 86-90.

BECKER, R.T., BOCKWINKEL, J., EBBIGHAUSEN, V., ABOUSSALAM, S.Z., ELHASSANI, A.,NÜBEL,H. 2004b. Lower and Middle Devonian stratigraphy and faunas at Bou Tserfine near Assa (Dra Valley, SW Morocco). Documents de l’Institut Scientifique, Rabat 19, 91-101.

BECKER, R.T., JANSEN, U., PLODOWSKI, G., SCHINDLER, E., ABOUSSALAM, S.Z. & WEDDIGE, K. 2004c. Devonian litho- and biostratigraphy of the Dra Valley - an overview. Documents de l’Institut Scientifique, Rabat 19, 5-20.

BONINO, E. & KIER, C. 2010. The Back to the Past Museum Guide to Trilobites. 494 pp. Back to the Past Museum, Puerto Morelos.

BRETT, C.E., ZAMBITO, J.J., SCHINDLER, E. & BECKER, R.T. 2012. Diagenetically-enhanced trilobite obrution deposits in concretionary limestones: The paradox of “rhythmic events beds”. Palaeogeography, Palaeoclimatology, Palaeoecology 367/368, 30-43.View article

BRONN, H. 1825. Ueber zwei neue Trilobiten-Arten zum Calymene-Geschlechte gehörig. Taschenbuch für die gesammte Mineralogie, 19(1), Zeitschrift für Mineralogie, Taschenbuch 1, 317-321.

BRUTON, D.L. & HAAS, W. 1997. Functional morphology of Phacopinae (Trilobita) and the mechanics of enrolment. Palaeontographica (A) 245, 1-43.

CAMPBELL, K.S.W. 1977. Trilobites of the Haragan, Bois d’Arc and Frisco formations (Early Devonian), Arbuckle Mountains region, Oklahoma. Oklahoma Geological Survey Bulletin 123, 1-227.

CHATTERTON, B.D.E. & GIBB, S.L. 2010. Latest Early to early Middle Devonian trilobites from the Erbenochile bed, Jbel Issoumour, southeastern Morocco. Journal of Paleontology 84, 1188-1205.View article

CHATTERTON, B.D.E., FORTEY, R.A., BRETT, K.D., GIBB, S.L. & MCKELLAR, R.C. 2006. Trilobites from the upper Lower to Middle Devonian Timrhanrhart Formation, Jbel Gara el Zguilma, southern Morocco. Palaeontographica Canadiana 25, 1-177.

CHLUPÁČ, I. 1971. New phacopid trilobites from the Devonian of Czechoslovakia. Časopis pro mineralogii a geologii 16, 255-261.

CHLUPÁČ, I. 1977. The phacopid trilobites of the Silurian and Devonian of Czechoslovakia. Rozpravy Ústředního ústavu geologického 43, 1-172.

CHLUPÁČ, I. & KUKAL, Z. 1986. Reflection of possible global Devonian events in the Barrandian area, C.S.S.R. In WALLISER, O.H. (ed.) Global Bio-events: a critical approach. Lecture Notes in Earth Sciences 8, 169-179.View article

CRÔNIER, C.&VIERSEN, A.P. VAN. 2008. The ‘Mur des douaniers’, an exceptionally well preserved Early Eifelian fossil site. Bulletin de la Société géologique de France 179, 89-95.View article

DALMAN, J.W. 1826. Om palaeaderna, eller de sa kallade trilobiterna. Kongliga Svenska Vetenskaps-Akademien Handlingar 1826, 113-294.

DORGAN, K.M., JUMARS, P.A., JOHNSON, B., BOUDREAU, B.P. & LANDIS, E. 2005. Burrow extension by crack propagation. Nature 433, 475.View article

DRESNAY, R. DU, HINDERMEYER, J., EMBERGER, A., CAIA, J., DESTOMBES, J. & HOLLARD, H. 1988. Carte géologique du Maroc, Todrha - Ma’der (Anti-Atlas oriental, zones axiale et péripherique Nord et Sud) - Echelle: 1/200 000. Notes et Mémoires du Service géologique du Maroc 243.

EBACH, M.C. & AHYONG, S.T. 2001. Phylogeny of the trilobite subgenus Acanthopyge (Lobopyge). Cladistics 17, 1-10.View article

EMMRICH, H.F. 1839. De trilobitis: dissertatio petrefactologica quam consensu et auctoritate amplissimi philosophorum ordinis in alma litterarum universitate Friderica Guilelma pro summis in philosophia honoribus. 56 pp. Nietackian, Berlin.

GHOBADI POUR,M. 2015. Name change for the Devonian trilobite Cultrops Struve, 1995. Paläontologische Zeitschrift 89, 669-670.View article

HAAS, W. 1968. Trilobiten aus dem Silur und Devon von Bithynien (NW-Türkei). Palaeontographica (A) 130, 60-207.

HAAS, W. 1969. Lower Devonian trilobites from central Nevada and northern Mexico. Journal of Paleontology 43, 641-659.

HAWLE, I. & CORDA, A.J.C. 1847. Prodrom einer Monographie der böhmischen Trilobiten. 176 pp. Calve, Prague.

HOLLOWAY, D.J. 2013. The trilobite Cirriticeps gen. nov. (Proetida, Tropidocoryphidae) from the Silurian of New South Wales. Memoirs of the Association of Australasian Palaeontologists 44, 53-63.

JANSEN, U., BECKER, G., PLODOWSKI, G., SCHINDLER, E., VOGEL, O. & WEDDIGE, K. 2004. Pragian and Emsian near Aouinet Torkoz (SW Dra Valley, Morocco). Documents de l’Institut Scientifique, Rabat 19, 75-84.

JELL, P.A. & ADRAIN, J.M. 2002. Available generic names for trilobites. Memoirs of the Queensland Museum 48, 331-553.

KHALDI, A.Y., CRÔNIER, C., HAINAUT, G., ABBACHE, A. & OUALI MEHADJI, A. 2016. A trilobite faunule from the Lower Devonian of the Saoura Valley, Algeria: biodiversity, morphological variability and palaeobiogeographical affinities. Geological Magazine 153, 357-387.View article

KIELAN, Z. 1954. Les trilobites mésodévoniens des monts de Sainte-Croix. Palaeontologia Polonica 6, 1-50.

KLUG, C., SCHULZ, H. DE BAETS, K. 2009. Red Devonian trilobites with green eyes from Morocco and the silicification of the trilobite exoskeleton. Acta Palaeontologica Polonica 54, 117-123.View article

LEMAÎTRE, D. 1952. La faune du Dévonien inférieur et moyen de la Souara et des abords de l’Erg El Djemel (Sud Oranais). Matériaux pour la Carte géologique de l’Algérie. 1re série, paléontologie 12, 1-170.

LESPÉRANCE, P.J. & SHEEHAN, P.M. 1987. Trilobites et brachiopods ashgilliens (Ordovicien supérieur) de l’«Assise» de Fosse, Bande de Sambre-Meuse (Belgique). Bulletin de l’Institut royal des Sciences naturelles de Belgique, Sciences de la Terre 57, 91-123.

LIEBERMAN, B.S. & KLOC, G.J. 1997. Evolutionary and biogeographic patterns in the Asteropyginae (Trilobita, Devonian) Delo, 1935. Bulletin of the American Museum of Natural History 232, 1-127.

MCKELLAR, R.C. & CHATTERTON, B.D.E. 2009. Early and Middle Devonian Phacopidae (Trilobita) of southern Morocco. Palaeontographica Canadiana 28, 1-109.

MORZADEC, P. 1969. Révision de Phacops (Phacops) potieri Bayle 1878 (Trilobites). Bulletin de la Société géologique de France (7) 10, 917-923.View article

MORZADEC, P. 1983. Trilobites du Dévonien (Emsien- Famennien) de la rade de Brest (Massif armoricain). Palaeontographica (A) 181, 103-184.

MORZADEC, P. 2001. Les trilobites Asteropyginae du Dévonien de l’Anti-Atlas (Maroc). Palaeontographica (A) 262, 53-85.View article

MORZADEC, P., MERGL, M., VILLARROEL, C., JANVIER, P. & RACHEBOEUF, P.R. 2015. Trilobites and inarticulate brachiopods from the Devonian Floresta Formation of Colombia: a review. Bulletin of Geosciences 90, 331-358.View article

MÜLLER, P. 2005. Revision der Gattung Scabrella (Trilobita; Unter-Devon). Geologica et Palaeontologica 39, 1-27.

ÖZDIKMEN, H. 2009. Nomenclatural changes for twenty trilobites genera. Munis Entomology & Zoology 4, 155-171.

PEK, I. & VANĚK, J. 1991. On some Silurian and Devonian trilobites of Bolivia. Acta Universitatis Palackianae Olomucensis Facultas Rerum Naturalium: Geographica-Geologica 103, 75-103.

PŘIBYL, A. & ERBEN, H.K. 1952. Über einige neue oder wenig bekannte Acanthopyginae (Tril.) des böhmischen und des deutschen Devons. Paläontologische Zeitschrift 26, 141-174.View article

SCHRAUT, G. 2000a. Trilobiten aus dem Unter-Devon des südöstlichen Anti-Atlas, Süd-Marokko. Senckenbergiana lethaea 79, 361-433.View article

SCHRAUT, G. 2000b. Eine neue Unterart von Phacops (Phacops) sparsinodosus Struve 1970 aus dem Mittel-Devon von Marokko. Senckenbergiana lethaea 80, 525-535.View article

SCHRAUT, G. & Feist, R. 2004. The Devonian styginid trilobite Paralejurus, with new data from Spain and Morocco. Journal of Paleontology 78, 709-722.View article

SCOTESE, C.R. 2014. Atlas of Devonian Paleogeographic Maps, PALEOMAP Atlas for ArcGIS, volume 4, The Late Paleozoic, Maps 65-72, Mollweide Projection. PALEOMAP Project, Evanston, IL.

SMEENK, Z. 1983. Devonian trilobites of the southern Cantabrian Mountains (northern Spain) with a systematic description of the Asteropyginae. Leidse Geologische Mededelingen 52, 383-511.

STRUVE, W. 1970. Beiträge zur Kenntnis der Phacopina (Trilobita), 7: Phacops-Arten aus dem Rheinischen Devon. 1. Senckenbergiana lethaea 51, 133-189.

STRUVE, W. 1972. Beiträge zur Kenntnis der Phacopina (Trilobita), 8: Phacops-Arten aus dem Rheinischen Devon. 2. Untergattungs-Zuweisung. Senckenbergiana lethaea 53, 383-403.

STRUVE, W. 1976. Beiträge zur Kenntnis der Phacopina (Trilobita), 9: Phacops (Omegops) n. sg. (Trilobita; Ober-Devon). Senckenbergiana lethaea 56, 429-451.

STRUVE, W. 1982. Beiträge zur Kenntnis der Phacopina (Trilobita), 10: Neue Untersuchungen über Geesops (Phacopinae; Unter- und Mittel-Devon). Senckenbergiana lethaea 63, 473-495.

STRUVE, W. 1995. Beiträge zur Kenntnis der Phacopina (Trilobita), 18: Die Riesen-Phacopiden aus dem Maider, SE-marokkanische Prä-Sahara. Senckenbergiana lethaea 75, 77-129.

THOMAS, A.T. & HOLLOWAY, D.J. 1988. Classification and phylogeny of the trilobite order Lichida. Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences 321(1205), 179-262.View article

VIERSEN, A.P. VAN 2006. New Middle Devonian trilobites from Vireux-Molhain (Ardennes, northern France). Senckenbergiana lethaea 86, 63-75.View article

VIERSEN, A.P. VAN 2007. Preliminary report of trilobites from the Hanonet Formation (Eifelian - Givetian transition), southern border of the Dinant Synclinorium, Belgium. Bulletin de l’Institut royal des Sciences naturelles de Belgique, Sciences de la Terre 77, 15-29.

VIERSEN, A.P. VAN 2008. First record of the odontopleurid trilobite Koneprusia from the lower Eifelian of Vireux-Molhain (northern France), with remarks on the associated trilobite fauna. Geologica Belgica 11, 83-91.

VIERSEN, A.P. VAN 2013. Latest Early to early Middle Devonian acastid trilobites from the eastern part of the Dinant Synclinorium, Belgium (Rhenohercynian Zone), 1-10. In HOLLOWAY, D.J. & LAURIE, J. (eds) Siluro-Devonian Studies. 2. Memoirs of the Association of Australasian Palaeontologists 44.

VIERSEN, A.P. VAN 2015. Trilobites du Dévonien des Ardennes (zone rhéno-hercynienne): biostratigraphie, évolution et événements. Fossiles, Revue française de Paléontologie 11, 5-24.

VIERSEN, A.P. VAN & HOLLAND, D. 2016. Morphological trends and new species of Cyphaspis (Trilobita, Otarioninae) in the Devonian of Morocco, Turkey, Germany and Belgium. Geologica Belgica 19, 251-271.View article

VIERSEN, A.P. VAN & PRESCHER, H. 2009. Trilobites from the Longlier Formation (Lower Devonian; Neufchâteau Synclinorium, southeast Belgium): first record of Pragian associated “Rhenish” and “Bohemian” assemblages from the Ardennes. Bulletin de l’Institut royal des Sciences naturelles de Belgique, Sciences de la Terre 79, 5-26.

VIERSEN, A.P. VAN, PRESCHER, H. & SAVELSBERGH, J. 2009. Description of two new trilobites from the Ahrdorf Formation (Middle Devonian) at the “Trilobitenfelder” of Gees, Eifel, Rhenish Mountains. Bulletin de l’Institut royal des Sciences naturelles de Belgique, Sciences de la Terre 79, 43-53.

VIERSEN, A.P. VAN, TAGHON, P. & MAGREAN, B. in press. The phacopid trilobites Austerops McKellar & Chatterton, 2009, Hottonops gen. nov. and Loreleiops gen. nov. from the Devonian of the Ardenno-Rhenish Mountains. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 283.

WRIGHT, A.J. & HAAS, W. 1990. A new Early Devonian spinose phacopid trilobite from Limekilns, New South Wales: morphology, affinities, taphonomy and palaeoenvironment. Records of the Australian Museum 42, 137-147.View article

WHITTINGTON, H.B. & KELLY, S.R.A. 1997. Morphological terms applied to Trilobita, 313-329. In KAESLER, R.L. (ed.) Treatise on invertebrate paleontology, Part O, Arthropoda 1. Trilobita, revised. Volume 1: Introduction, Order Agnostida, Order Redlichiida. Geological Society of America, Boulder, Colorado and University of Kansas Press, Lawrence, Kansas.