Sponge growth on the Cenomanian carbonate shelves of the Carpathian Basin: a record from spicule-rich turbidites


Authors: Bąk M, Gorny Z, Bąk K

Published in: Bulletin of Geosciences, volume 90, issue 3; pages: 651 - 666; Received 14 November 2014; Accepted in revised form 8 July 2015; Online 8 September 2015

Keywords: demosponge, spiculite, turbidite, Silesian Basin, Outer Carpathians, Cenomanian,

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The composition and features of numerous bioclasts found in the sandy/silty fraction of turbidite layers in the Middle-Upper Cenomanian deep-water sediments of the Silesian Nappe in the Outer Carpathians, dominated by sponge spicules, suggest that they originated from the growth and destruction of sponge assemblages formed on shelves of the northern margin of the Silesian Basin. The morphological types of spicules, by comparison with fossil and living species, lead to their attribution predominantly to three orders: Lithistida, Hadromerida and Astrophorida. Sponges lived in association with other benthic fauna such as calcareous benthic foraminifers and echinoids. Many sponge spicules were calcified in their primary environment before their redeposition by turbiditic flows. Favourable conditions for development of sponge assemblages occurred during the Cenomanian transgressions, being related to balanced delivery of sediment particles trapped by sponges and waters enriched with nutrients. The amount of nutrients absorbed by sponges is related to the dissolved silica content that originated from two sources: 1) terrestrial chemical weathering on the low-gradient plains of the shelves and 2) opal grains, such as siliceous plankton, suspended in the water column during bloom periods forced by seasonal upwelling.


ALEXANDROWICZ, S.W. 1973. Gaize-type sediments in the Carpathian flysch. Neues Jahrbuch für Geologie und Paläontologie, Monatschefte 1, 1-17.

BAVESTRELLO, G., CATTANEO-VIETTI, R., CERRANO, C. & SARA, M. 1996. Spicule dissolution in living Tethya omanensis (Porifera: Demospongiae) from a tropical cave. Bulletin of Marine Science 58, 598-601.

BAVESTRELLO, G., BENATTI, U., CATTANEO-VIETTI, R., CERRANO, C. & GIOVINE, M. 2003. Sponge cell reactivity to various forms of silica. Microscopy Research and Technique 62, 327-335.View article

BĄK, K. 2006. Sedimentological, geochemical and microfaunal responses to environmental changes around the Cenomanian-Turonian boundary in the Outer Carpathian Basin; a record from the Subsilesian Nappe, Poland. Palaeogeography, Palaeoclimatology, Palaeoecology 23, 335-358.View article

BĄK, K. 2007a. Deep-water facies succession around the Cenomanian-Turonian boundary in the Outer Carpathian Basin: Sedimentary, biotic and chemical records in the Silesian Nappe, Poland. Palaeogeography, Palaeoclimatology, Palaeoecology 248, 255-290.View article

BĄK, K. 2007b. Organic-rich and manganese sedimentation during the Cenomanian-Turonian boundary event in the Outer Carpathian Basin, a new record from the Skole Nappe, Poland. Palaeogeography, Palaeoclimatology, Palaeoecology 256, 21-46.View article

BĄK, K. 2007c. Environmental changes during the Cenomanian-Turonian boundary event in the Outer Carpathian basins: a synthesis of data from various tectonic-facies units. Annales Societatis Geologorum Poloniae 77, 171-191.

BĄK, K., BĄK, M. & PAUL, Z. 2001. Barnasiówka Radiolarian Shale Formation - a new lithostratigraphic unit in the Upper Cenomanian-lowermost Turonian of the Polish Outer Carpathians. Annales Societatis Geologorum Poloniae 71, 75-103.

BĄK, M. 2000. Radiolaria from the Upper Cenomanian-Lower Turonian deposits of the Silesian Unit (Polish Flysch Carpathians). Geologica Carpathica 51, 309-324.

BĄK, M. 2004. Radiolarian biostratigraphy of the Upper Cenomanian-Lower Turonian deposits in the Subsilesian Nappe (Outer Western Carpathians). Geologica Carpathica 55, 239-250.

BĄK, M. 2011. Tethyan radiolarians at the Cenomanian-Turonian Anoxic Event from the Apennines (Umbria-Marche) and the Outer Carpathians: Palaeoecological and palaeoenvironmental implications. Studia Geologica Polonica 134, 7-279.

BĄK, M., BĄK, K. & CIUREJ, A. 2005. Mid-Cretaceous spicule-rich flysch deposits in the Silesian Nappe of the Polish Outer Carpathians; radiolarian and foraminiferal biostratigraphy. Geological Quarterly 49, 275-290.

BĄK, M., BĄK, K. & CIUREJ, A. 2011. Palaeoenvironmental signal from the microfossils record in the Mikuszowice Cherts of the Silesian Nappe, Polish Outer Carpathians, 15-25. In BĄK, M., KAMINSKI, M.A. & WAŚKOWSKA, A. (eds) Integrating Microfossil Records from the Oceans and Epicontinental Seas. Grzybowski Foundation Special Publication 17. 144 pp. Grzybowski Foundation, Kraków.

BĄK, M., BĄK, K., GÓRNY, Z., STOŻEK, B. & WOLSKA, A. 2015. Evidence of bacteriogenic iron and manganese oxyhydroxides in Albian-Cenomanian marine sediments of the Carpathian realm (Poland). Annales Societatis Geologorum Poloniae 85(2), 371-385. BIRKENMAJER, K. 1986. Stages of structural evolution of the Pieniny Klippen Belt, Carpathians. Studia Geologica Polonica 88, 7-32.

BRYNDAL, T. 2014. A method for identification of small Carpathian catchments more prone to flash flood generation. Based on the example of south-eastern part of the Polish Carpathians. Carpathian Journal of Earth and Environmental Sciences 9, 109-122.

BURTAN, J. (ed.) 1964. Detailed Geological Map of Poland in scale 1:50,000 (without Quaternary sediments); the Carpathian and Fore-Carpathian area: Myślenice sheet. Wydawnictwa Geologiczne, Warszawa.

BURTAN, J. & SZYMAKOWSKA, F. (eds) 1964. Detailed Geological Map of Poland in scale 1:50,000 (without Quaternary sediments); the Carpathian and Fore-Carpathian area: Osielec sheet. Wydawnictwa Geologiczne, Warszawa.

CARTER, H.J. 1871. On fossil sponge-spicules of the Greensland compared with those of existing species. Annals and Magazine of Natural History, Series 4, 7(38), 112-141.

DEFRETIN-LEFRANC, S. 1960. Contribution a l’étude des spongiaires siliceux du Crétacé supérieur du Nord de la France. Theses a la Faculte des Sciences de Lille 1958, 1-178.

DUARTE, L.V., KRAUTTER, M. & SOARES, A.F. 2001. Bioconstructions a spongiaires siliceux dans le Lias terminal du Bassin lusitanien (Portugal): stratigraphie, sédimentologie et signification paléogéographique. Bulletin de la Société géologique de France 172, 637-646.View article

DUBICKA, Z. & PERYT, D. 2014. Classification and evolutionary interpretation of late Turonian-early Campanian Gavelinella and Stensioeina (Gavelinellidae, benthic foraminifera) from western Ukraine. Journal of Foraminiferal Research 44, 151-176.View article

FROGET, C. 1976. Obsérvations sur l’altération de la silice et des silicates au cours de la lithification carbonatée (Région Siculo-Tunisienne). Géologie Méditerréenne 111, 219- 226.

GAMMON, P.R., JAMES, N.P. & PISERA, A. 2000. Eocene spiculites and spongolites in southwestern Australia: not deep, not polar, but shallow and warm. Geology 28, 855-858.View article

GAWOR-BIEDOWA, E. 1972. The Albian, Cenomanian and Turonian foraminifers of Poland and their stratigraphic importance. Acta Paleontologica Polonica 17, 3-165.

GOLONKA, J., OSZCZYPKO, N. & ŚLĄCZKA, A. 2000. Late Carboniferous-Neogene geodynamic evolution and paleogeography of the circum-Carpathians region and adjacent areas. Annales Societatis Geologorum Poloniae 70, 107-136.

GRÄFE, K.-U. 2005. Late Cretaceous benthic foraminifers from Basque-Cantabrian Basin, Northern Spain. Iberian Geology 31, 277-298.

GUTOWSKI, J., POPADJUK, I.V. & OLSZEWSKA, B. 2005. Late Jurassic-earliest Cretaceous evolution of the epicontinental sedimentary basin of southeastern Poland and Western Ukraine. Geological Quarterly 49, 45-52.

HAMMES, U. 1995. Initiation and development of small-scale sponge mud-mounds, Late Jurassic, Southern Franconian Alb, Germany, 335-357. In MONTY, C.L.V., BOSENCE, D.W.J., BRIDGES, P.H. & PRATT, B.R. (eds) Carbonate mud-mounds: their origin and evolution. International Associations of Sedimentologists Special Publications 23. Blackwell, Oxford.

HAQ, B.U. 2014. Cretaceous isostasy revisited. Global and Planetary Change 113, 44-58.View article

HART, M.U. 1980. A water depth model for the evolution of the planktonic foraminifera. Nature 286, 252-254.View article

HART, M.B. & SWIECICKI, A. 1988. The genus Gavelinella Brotzen, 1942, in the Cretaceous strata of the U.K. Revue Paléobiologie vol. spéc. 2, 289-294.

HELLER, I. 1975. Microbiostratigraphy of the Cretaceous deposits in the southern part of the Łódź Synklinorium (Central Poland). Annales Societatis Geologorum Poloniae 55, 233-254.

HENTSCHEL, U., SCHMID, M., WAGNER, M., FIESELER, L., GERNERT, C. & HACKER, J. 2001. Isolation and phylogenetic analysis of bacteria with antimicrobial activities from the Mediterranean sponges Aplysina aerophoba and Aplysina cavernicola. FEMS Microbiology Ecology 35, 305-312.View article

HINDE, G.J. 1883. Catalogue of the fossil sponges in the Geological Department of the British Museum. 248 pp. British Museum (Natural History), London.

HOOPER, J.N.A. & VAN SOEST, R.W.M. 2002. Order Astrophorida Sollas, 186-195. In HOOPER, J.N.A. & VAN SOEST, R.W.M. (eds) Systema Porifera. Kluwer Academics/Plenum Press, New York.

HOOPER, J.N.A., VAN SOEST, R.W.M. & PISERA, A. 2011. Phylum Porifera Grant, 1826, 13-18. In ZHANG, Z.-Q. (ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148.

HRADECKÁ, L. 1993. Gavelinella Brotzen, 1942 and Lingulogavelinella Malapris, 1969 (Foraminifera) from the Bohemian Basin. Sborník geologických věd, Paleontologie 33, 79-96.

HRADECKÁ, L. & ŠVÁBENICKÁ, L. 1995. Foraminifera and calcareous nannoplankton assemblages from the Cenomanian-Turonian boundary interval of the Knoviz section, Bohemian Cretaceous Basin. Geologica Carpathica 46, 267-276.

HURCEWICZ, H. 1966. Siliceous sponges from the Upper Cretaceous of Poland. Part I. Tetraxonia. Acta Paleontologica Polonica 11, 15-119.

HURCEWICZ, H. 1968. Siliceous sponges from the Upper Cretaceous of Poland, Part II, Monaxonia and Triaxonia. Acta Palaeonologica Polonica 13, 3-96.

JAŚKOWIAK-SCHOENEICHOWA, M. & KRASSOWSKA, A. 1988. Palaeothickness, lithofacies and palaeotectonics of the epicontinental Upper Cretaceous in Poland. Kwartalnik Geologiczny 32, 177-198.

KAUFFMAN, E.G., HERM, D., JOHNSON, C.C., HARRIES, P. & EFLING, R.H. 2000. The ecology of Cenomanian lithistid sponge frameworks, Regensburg area, Germany. Lethaia 33, 214-235.View article

KOSZARSKI, L. & ŚLĄCZKA, A. 1973. Outer (flysch) Carpathians, Lower Cretaceous, 492-495. In POŻARYSKI, W. (ed.) Geology of Poland. Instytut Geologiczny, Warszawa.

KRAUTTER, M. 1997. Aspekte zur Paläökologie postpaläozoischer Kieselschwämme. Profil 11, 199-324.

KRAUTTER, M., CONWAY, K.W., BARRIE, J.V. & NEUWEILER, M. 2001. Discovery of a “living dinosaur”: globally unique modern Hexactinellid sponge reefs off British Columbia, Canada. Facies 44, 265-282.View article

KSIĄŻKIEWICZ, M. 1951. Explanations to the Wadowice sheet of the Detailed Geological Map of Poland in scale 1:50 000. 283 pp. Państwowy Instytut Geologiczny, Warszawa.

KSIĄŻKIEWICZ, M. 1956. Geology of the Northern Carpathians. Geologische Rundschau 45, 396-411.View article

KSIĄŻKIEWICZ, M. (ed.) 1962. Geological Atlas of Poland; Stratigraphic and Facial Problems, vol. 13. Cretaceous and Older Paleogene in the Polish Outer Carpathians. 20 maps, 20 pp. Instytut Geologiczny, Wydawnictwa Geologiczne, Warszawa.

KSIĄŻKIEWICZ, M. 1977. Hypothesis of plate tectonics and the origin of the Carpathians. Annales Societatis Geologorum Poloniae 47, 329-353.

KUTEK, J. 1994. Jurassic tectonic events in south-eastern cratonic Poland. Acta Geologica Polonica 44, 167-221.

LAND, L.S. 1976. Early dissolution of sponge spicules from reef sediments, North Jamaica. Journal of Sedimentary Petrology 46, 967-969.

LAUBENFELS, M.W. DE 1955. Archaeocyatha and Porifera, 1-113. In MOORE, R.C. (ed.) Treatise on Invertebrate Paleontology. Part E. Geological Society of America & University of Kansas Press, Boulder & Lawrence.

LECKIE, R.M. 1987. Paleoecology of mid-Cretaceous planktonic foraminifera: a comparison of open ocean and epicontinental sea assemblages. Micropaleontology 33, 164-176.View article

LÉVI, C. 1991. Lithistid sponges from the Norkfolk Rise. Recent and Mesozoic genera, 72- 82. In REITNER, J. & KEUPP, H. (eds) Fossil and recent sponges. Springer Verlag, Berlin.View article

ŁUKOWIAK, M. 2015. Late Eocene siliceous sponge fauna of southern Australia: reconstruction based on loose spicules record. Zootaxa 3917(1), 001-065.View article

MALDONADO, M., CARMONA, M.C., URIZ, M.J. & CRUZADO, A. 1999. Decline in Mesozoic reef-building sponges explained by silicon limitation. Nature 401, 785-788.View article

MAREK, S. 1988. Palaeothickness, lithofacies and palaeotectonics of the epicontinental Lower Cretaceous in Poland. Kwartalnik Geologiczny 32, 157-174.

MILLER, K.G., KOMINZ, M.A., BROWNING, J.V., WRIGHT, J.D., MOUNTAIN, G.S., KATZ, M.E., SUGARMAN, P.J., CRAMER, B.S., CHRISTIE-BLICK, N. & PEKAR, S.F. 2005. The Phanerozoic record of global sea-level change. Science 310, 1293-1298.View article

MORET, L. 1926. Contribution a l’étude des spongiaires siliceux du Crétacé supérieur français. Mémoire de la Société géologique de France, nouvelle série 5, 1-314.

MORYC, W. 1997. The Lower Cretaceous in the pre-Miocene substratum of the southern part of the Carpathian Foredeep in Poland. Annales Societatis Geologorum Poloniae 67, 287-296.

NESTLER, H. 1961. Spongien aus der weissen Schreibkreide (unter Maastricht) der Insel Rügen (Ostsee). Paläontologische Abhandlungen 1, 1-70.

NEUMANN, A.C., KOFOED, J.W. & KELLER, G.H. 1977. Lithoherms in the Straits of Florida. Geology 1, 4-10.View article

NEUWAILER, F. & BURDIGE, D.J. 2005. The modern calcifying sponge Spheciospongia vesparium (Lamarck, 1815), Great Bahama Bank: Implications for ancient sponge mud-mounds. Sedimentary Geology 175, 89-98.View article

OKOŃSKI, S., GÓRNY, Z., BĄK, M. & BĄK, K. 2014. Lithistid spicules in the sediments of the Turonian Variegated Shale in the Silesian Nappe, Polish Outer Carpathians. Geology, Geophysics & Environment 40, 33-48.View article

OLSZEWSKA-NEJBERT, D. & ŚWIERCZEWSKA-GŁADYSZ, E. 2011. Campanian (Late Cretaceous) hexactinellid sponges from the white chalk of Mielnik (Eastern Poland). Acta Geologica Polonica 61(4), 38-417.

OLSZEWSKA-NEJBERT, D. & ŚWIERCZEWSKA-GŁADYSZ, E. 2013. Cenomanian (Late Cretaceous) siliceous sponges from Nezvys’ko and Rakovets’ (Pokuttian Podillia, western Ukraine). Cretaceous Research 43, 116-144.View article

OSZCZYPKO, N. 2004. The structural position and tectonosedimentary evolution of the Polish Outer Carpathians. Przegląd Geologiczny 52, 780-791.

PERYT, D. 1983. Mid-Cretaceous microbiostratigraphy and foraminifers of the NE margins of the Świętokrzyskie (Holy Cross) Mts., Poland. Acta Paleontologica Polonica 28, 417-466.

PISERA, A. 1997. Upper Jurassic siliceous sponges from the Swabian Alb: taxonomy and paleoecology. Palaeontologia Polonica 57, 1-214.

PISERA, A. 1999. Post-Palaeozoic history of the siliceous sponges with rigid skeleton. Memoires of the Queensland Museum 44, 463-472.

PISERA, A. 2006. Palaeontology of sponges - a review. Canadian Journal of Zoology 84, 242-261.View article

PISERA, A., CACHAO, M. & SILVA, C.M. 2006. Siliceous sponge spicules from the Miocene Mem Moniz marls (Portugal) and their environmental significance. Rivista Italiana di Paleontologia e Stratigrafia 112(2), 287-299.

PISERA, A. & LÉVI, C. 2002a. Family Corallistidae Sollas, 312-320. In HOOPER, J.N.A. & VAN SOEST, R.W.M. (eds) Systema Porifera. Kluwer Academics/Plenum Press, New York.

PISERA, A. & LÉVI, C. 2002b. Family Phymaraphiniidae Schrammen, 380-383. In HOOPER, J.N.A. & VAN SOEST, R.W.M. (eds) Systema Porifera. Kluwer Academics/Plenum Press, New York.

PISERA, A. & LÉVI, C. 2002c. Family Phymatellidae Schrammen, 366-373. In HOOPER, J.N.A. & VAN SOEST, R.W.M. (eds) Systema Porifera. Kluwer Academics/Plenum Press, New York.

PISERA, A. & LÉVI, C. 2002d. Family Theonelleidae Lendenfeld, 327-337. In HOOPER, J.N.A. & VAN SOEST, R.W.M. (eds) Systema Porifera. Kluwer Academics/Plenum Press, New York.

PISERA, A., SATIR, M., GRUSZCZYŃSKI, M., HOFFMAN, A. & MAŁKOWSKI, K. 1992. Variation in δ13C and δ18O in late Jurassic carbonates, Submediterranean Province, Europe. Annales Societarum Geologorum Poloniae 62, 141-147.

PRATT, B.R., BOURQUE, P.A. & GIGNAC, H. 1986. Sponge-constructed stromatactis mud mounds, Silurian of Gaspe, Quebec; discussion and reply. Journal of Sedimentary Research 56, 459-463.View article

REID, R.E.H. 1958. A monograph of the Upper Cretaceous Hexactinellida of Great Britain and Northern Ireland, part II, 1-26. Palaeontographical Society, London.

REID, R.E.H. 1962. Sponges and the Chalk Rock. Geological Magazine 99, 273-278.View article

REID, R.E.H. 2004. Mesozoic and Cenozoic choristid demosponges, 175-198. In RIGBY, J.K., FINKS, R.M. & REID, R.E.H. (eds) Treatise on Invertebrate Paleontology. Part E, Vol. 3. Porifera. Geological Society of America & University of Kansas Press, Boulder & Lawrence.

REITNER, J. 1987. Mikrofazielle, paläekologische und paläogeographische analyse ausgewählter vorkommen flachmariner Karbonate im Basko-Kantabrischen Strike Slip Fault-Becken-System (Nordspanien) an der Wende von der Unterkreide zur Oberkreide. Documenta naturae 40, 1-239.

REITNER, J. 1993. Modern cryptic microbialite-metazoan facies from Lizard Island (Great Barrier Reef, Australia) - formation and concepts. Facies 29, 3-40.View article

REITNER, J. & KEUPP, H. 1991. The fossil record of the haplosclerid excavating sponge Aka de Laubenfels, 102-120. In REITNER, J. & KEUPP, H. (eds) Fossil and Recent Sponges. Springer Verlag, Berlin.

REITNER, J., NEUWEILER, F. & GAUTRET, P. 1995. Modern and fossil automicrites: implications for mud-mound genesis. Facies 32, 4-17.View article

RICE, A.L., THURSTON, M.H. & NEW, A.L. 1990. Dense aggregations of a hexactinellid sponge, Pheronema carpenteri, in the Porcupine Seabight (northeast Atlantic Ocean) and possible causes. Progress in Oceanography 24, 179-206.View article

ROEMER, F.A. 1864. Die Spongitarien des norddeutschen Kreidegebirges. Palaeontographica 13, 1-63.

ROTHWELL, R.G. 1989. Minerals and mineraloids in marine sediments: An optical identification guide. 279 pp. Elsevier Applied Science, London.View article

SCHRAMMEN, A. 1910. Die Kieselspongien der oberen Kreide von Nordwestdeutschland. I. Teil. Tetraxonia, Monaxonia und Silicea incertae sedis. Palaeontographica 5, Supplement, 1-175.

SCHRAMMEN, A. 1910-1912. Die Kieselspongien der oberen Kreide von Nordwestdeutschland. Palaeontographica 5, Supplement, 1-385.

SUJKOWSKI, Z. 1933. Sur certains spongiolithes de la Tatra et des Karpates. Sprawozdania - Państwowy Instytut Geologiczny 7, 712-733.

ŚWIERCZEWSKA-GŁADYSZ, E. 2006. Late Cretaceous siliceous sponges from the Middle Vistula River Valley (Central Poland) and their palaeoecological significance. Annales Societatis Geologorum Poloniae 76, 227-296.

ŚWIERCZEWSKA-GŁADYSZ, E. 2012. Hexactinellid sponge assemblages across the Campanian-Maastrichtian boundary in the Middle Vistula River section, central Poland. Acta Geologica Polonica 62(4), 561-580.View article

TERMIER, G. & TERMIER, H. 1980. Spongiaires du Crétacé Moyen. Cretaceous Research 2, 427-433.View article

TREST’JAN, G.N. 1972. New glass sponges (Hyalospongiae) from the Upper Cretaceous of the Pridnestrovie. Paleontologicheskii zhurnal 6, 171-179. [in Russian]

UNRUG, R. 1959. On the sedimentation of the Lgota beds. Rocznik Polskiego Towarzystwa Geologicznego 29, 197-225.

UNRUG, R. 1977. Ancient deep-sea traction currents deposits in the Lgota beds (Albian) of the Carpathian Flysch. Rocznik Polskiego Towarzystwa Geologicznego 47, 355-370.

ULBRICH, H. 1974. Die Spongien der Ilsenburg-Entwicklung (Oberes Unter-Campan) der Subherzynen Kreidemulde. Freiberger Forschungshefte C 291, 1-121.

URIZ, M.J. 2002. Family Geodiidae Gray, 1867, 134-140. In HOOPER, J.N.A. & VAN SOEST, R.W.M. (eds) Systema Porifera. Kluwer Academics/Plenum Press, New York.

VACELET, J. 1975. Étude en microscopie électronique de l’association entre bactéries et spongiaires du genre Verongia (Dictyoceratida). Journal of Microscopy 23, 271-288.

VAN WEERING, T.C.E., DE HAAS, H., DE STIGTER, H.C., LYKKE-ANDERSEN, H. & KOUVAEV, I. 2003. Structure and development of Giant carbonate mounds at the SW and SE Rockall Trough margin. Marine Geology 198, 67-83.View article

VOIGT, S., AURAG, A., LEIS, F. & KAPLAN, U. 2007. Late Cenomanian to Middle Turonian high-resolution carbon isotope stratigraphy: New data from the Münsterland Cretaceous Basin, Germany. Earth and Planetary Science Letters 253, 196-210.View article

WARNKE, K. 1995. Calcification processes of siliceous sponges in Viséan limestones (counties Sligo and Leitrim, Northwestern Ireland). Facies 33, 215-228.View article

WHITNEY, F., CONWAY, K., THOMSON, R., BARRIEB, V., KRAUTTER, M. & MUNGOV, G. 2005. Oceanographic habitat of sponge reefs on the Western Canadian continental shelf. Continental Shelf Research 25, 211-226.View article

WIEDENMAYER, F. 1980. Siliceous sponges; Development through time, 55-85. In HARTMAN, W.D., WENDT, J.W. & WIEDENMAYER, F. (eds) Living and fossil sponges. Notes for a short course, Sedimenta 8.

WIEDENMAYER, F. 1994. Contribution of the knowledge of post-Paleozoic neritic and archibenthal sponges (Porifera). Schweizerische Paläontologische Mitteilungen 116, 1-147.