The effaced Ordovician trilobite Svobodapeltis and the earliest history of illaenimorph trilobites in the Ordovician of the Prague Basin (Bohemia)

 

Authors: Mergl M, Kozák V

Published in: Bulletin of Geosciences, volume 91, issue 1; pages: 155 - 168; Received 23 April 2014; Accepted in revised form 27 January 2016; Online 15 March 2016

Keywords: trilobites, Illaenidae, Svobodapeltis, Hemibarrandia, Darriwilian, Ordovician, Czech Republic,

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Abstract

Revision of the Middle Ordovician (Darriwilian) illaenimorph trilobite Svobodapeltis avus (Holub, 1908) results in a substantially emended diagnosis of the genus Svobodapeltis Šnajdr, 1957 and its assignment to the Family Hemibarrandiidae Courtessole & Pillet, 1975. The general illaeniform morphology of Svobodapeltis and its relative Hemibarrandia Prantl & Přibyl, 1949, with effaced cephalon and pygidium, the connective suture on the cephalic doublure indicating the presence of a rostral plate, no articulating furrows in the axis, and the fulcrate thorax are in distinct contrast with the presence of pleural furrows on all thoracic segments. The glabella that is defined anteriorly, the presence of border furrows on the cephalon, lack of the lunette, and furrowed segments are regarded as convincing evidence for separation of the Hemibarrandiidae from other illaenoids. The illaenid Caudillaenus advena (Barrande, 1872), a specimen of which was previously assigned to S. avus that influenced the concept of the genus, is redescribed together with its ancestor C. aff. advena.

References

ANGELIN, N.P. 1854. Palaeontologia Scandinavica, Iconographia Crustacea Formationis Transitionis. Fasc. II, 21-92. Weigel, Lund.

BARRANDE, J. 1846. Notice préliminaire sur le Systeme silurien et les trilobites de Boheme. vi + 97 pp. Hirschfeld, Leipzig.

BARRANDE, J. 1872. Systeme silurien du Centre de la Boheme. Iere partie. Recherches paléontologiques. Supplément au Vol. 1. Trilobites, Crustacés divers et Poisson. 647 pp. Chez l’auteur et éditeur, Imprimerie de Charles Bellmann, Prag.

BRUTHANSOVÁ, J. 2003. The trilobite Family Illaenidae Hawle et Corda, 1847 from the Ordovician of the Prague Basin (Czech Republic). Transactions of the Royal Society of Edinburgh, Earth Sciences 93, 167-190.

BRUTON, D.L. 1968. The trilobite genus Panderia from the Ordovician of Scandinavia and the Baltic areas. Norsk Geologisk Tidsskrift 48, 1-53.

BURMEISTER, H. 1843. Die Organisation der Trilobiten, aus ihren lebenden Verwandten entwickelt; nebst einer systematischen Uebersicht aller zeither beschriebenen Arten. xii + 148 pp. Georg Reimer, Berlin.

COURTESSOLE, R. & PILLET, J. 1975. Contribution a l’étude des faunes trilobitiques de l’Ordovicien inférieur de la Montagne Noire. Les Eulominae et les Nileidae. Annales de la Société géologique du Nord 95, 251-272.

DALMAN, J.W. 1827. Om Palaeaderna eller de sa kallade trilobiterna. Kongliga Svenska Vetenskap-Akademiens Handlingar 1826, 113-162, 226-294.

DOUBRAVA, M. & VOKÁČ, V. 2004. Ectillaenus aff. advena (Barrande, 1872) (Trilobita) from the Arenigian (Klabava Formation, Lower Ordovician) of the Prague Basin (Barrandian, Bohemia). Journal of the Czech Geological Society 49(3-4), 125-126.

FORTEY, R.A. & BRUTON, D.L. 2013. Lower Ordovician trilobites of the Kirtonryggen Formation, Spitsbergen. Fossils and Strata 59, 1-116.View article

FORTEY, R.A. & OWENS, R.M. 1987. The Arenig Series in South Wales: Stratigraphy and Palaeontology. Bulletin of the British Museum (Natural History), Geology 41(3), 169-307.

HAVLÍČEK, V. 1998. Ordovician, 41-79. In CHLUPÁČ, I., HAVLÍČEK, V., KŘÍŽ, J., KUKAL, Z., & ŠTORCH, P. Palaeozoic of the Barrandian (Cambrian to Devonian). 183 pp. Czech Geological Survey, Prague.

HAVLÍČEK, V. & VANĚK, J. 1966. The biostratigraphy of the Ordovician of Bohemia. Sborník geologických věd, Paleontologie 8, 7-69.

HAWLE, I. & CORDA, A.J.C. 1847. Prodrom einer Monographie der böhmischen Trilobiten. 176 pp. J.G. Calve’sche Buchhandlung, Prague.

HOLLOWAY, D.J. 2007. The trilobite Protostygina and the composition of the Styginae, with two new genera. Paläontologische Zeitschrift 81(1), 1-16.View article

HOLUB, K. 1908. Příspěvek ku poznání fauny pásma Ddγ. Rozpravy České akademie věd a umění, Třída 2 17(10), 1-18.

HORNÝ, R. & BASTL, F. 1970. Type Specimens of Fossils in the National Museum Prague, volume 1, Trilobita. 354 pp. Museum of Natural History, Prague.

INGHAM, J.K. & TRIPP, R.P. 1991. The trilobite fauna of the Middle Ordovician Doularg Formation of the Girvan District, Scotland, and its palaeoenvironmental significance. Transactions of the Royal Society of Edinburgh, Earth Sciences 82, 27-54.View article

JAANUSSON, V. 1954. Zur Morphologie und Taxonomie der Illaeniden. Arkiv för Mineralogi och Geologi 1(20), 545-583.

JAANUSSON, V. 1957. Unterordovizische Illaeniden aus Skandinavien. Mit Bemerkungen über die Korrelation des Unterordoviziums. Bulletin of the Geological Institutions of the University of Uppsala 37(1-2), 79-165.

JAANUSSON, V. 1959. Suborder Illaenina, O365-O376. In MOORE, R.C. (ed.) Treatise on Invertebrate Paleontology. Part O, Arthropoda 1. 560 pp. Geological Society of America & University of Kansas Press, Lawrence.

KLOUČEK, C. 1916. O vrstvách dγ, jich trilobitech a nalezištích. Rozpravy České akademie věd a umění, Třída 2 25(39), 1-20.

KOBAYASHI, T. 1935. The Cambro-Ordovician Formations and Faunas of South Chosen. Paleontology III. Cambrian faunas of South Chosen. Journal of the Faculty of Science, Imperial University of Tokyo, Section II - Geology, Mineralogy, Geography, Seismology 4, 49-344.

KRAFT, J. 1972. Type specimens of fossils in the West-Bohemian Museum in Plzeň and in the Regional Museum of Dr. B. Horák in Rokycany. Part I - Trilobita. Folia Musei rerum naturalium Bohemiae occidentalis, Geologica 1, 1-11.

KRAFT, J. & KRAFT, P. 1999. Graptolite biostratigraphy of the Lower and Middle Ordovician of Bohemia. Acta Universitatis Carolinae, Geologica 33(1- 2), 33-36.

LINNAEUS, C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentis, synonymis, locis. Tomus I. Editio decima, reformata. 824 pp. Laurentii Salvii, Holmiae.View article

LU, Y.H. & ZHOU, T.R. 1990. Trilobites across the Cambrian-Ordovician Boundary of the transitional region of Sandu, southeastern Guizhou. Palaeontologia Cathayana 5, 1-84.View article

MERGL, M. 1979. Eccoptochile aff. clavigera (Beyrich, 1845) and Pliomerops lindaueri (Barrande, 1846) (Trilobita) from the Lower Ordovician of Bohemia. Věstník Ústředního ústavu geologického 54(5), 175-177.

MERGL, M. 1991a. New Lower Ordovician (Arenig) trilobite assemblages in Bohemia. Časopis pro mineralogii a geologii 36(4), 193-203.

MERGL, M. 1991b. Arenig (Lower Ordovician) orthide brachiopods from Prague Basin, Bohemia. Časopis pro mineralogii a geologii 36(1), 1-13.

MERGL, M. 1994. Trilobite fauna from the Třenice Formation (Tremadoc) in Central Bohemia. Folia Musei rerum naturalium Bohemiae occidentalis, Geologica 39, 1-31.

MERGL, M. 2006. Tremadocian trilobites of the Prague Basin, Czech Republic. Acta Musei nationalis Pragae, Series B - Historia Naturalis 62(1-2), 1-70.

MERGL, M., FATKA, O. & BUDIL, P. 2007. Lower and early Middle Ordovician trilobite associations of the Prague Basin (Perunica, Czech Republic). Acta Palaeontologica Sinica 46 (suppl.), 320-327.

MOBERG, J.C. & SEGERBERG, C.O. 1906. Bildrag till kännedomen om Ceratopygeregionen med särskild hänsyn till dess utveckling i Fogelsongstrakten. Meddelande Lunds Geologiska Fältklubb 2, 1-113.

NOVÁK, O. 1884. Studien an Hypostomen böhmischer Trilobiten, Nro. II. Sitzungsberichte der Königl. Böhmischen Geselsschaft der Wissenschaften in Prag, 1-20.

NOVÁK, O.P. 1918. Trilobiti pásma D-d1γ z okolí pražského. Rukopisná studie, k tisku upravil a doplňky opatřil Jaroslav Perner. Palaeontographica Bohemiae 9, 1-28.

NICHOLSON, H.A. & ETHERIDGE, R. 1876. A monograph of the Silurian fossils of Girvan in Ayshire with special reference to those contained in ’Gray Collection’, Volume 1(2). i-vi, 137-236. William Blackwood & Sons, Edinburgh & London.

PEK, I. & VANĚK, J. 1989. Index of Bohemian Trilobites. 168 pp. Krajské vlastivědné muzeum Olomouc, Olomouc.

POULSEN, O. 1959. Suborder Illaenina, O356-O359. In MOORE, R.C. (ed.) Treatise on Invertebrate Paleontology. Part O, Arthropoda 1. 560 pp. Geological Society of America & University of Kansas Press, Lawrence.

PRANTL, F. & PŘIBYL, A. 1949. On the genus Symphysurus Goldfuss and allied forms from the Ordovician of Bohemia (Trilobitae). Věstník Královské české společnosti nauk, Třída matematicko-přírodovědecká 1948(12), 1-16.

PŘIBYL, A. 1953. Seznam českých trilobitových rodů. Knihovna Ústředního ústavu geologického 25, 1-80.

RÁBANO, I. 1990. Trilobites del Ordovícíco Medio del sector meridional de la zona Centroiberica Espanola. Boletín geológico y minero, publicaciones especiales I, VII-XII, 1-233.

RÁBANO, I., GUTIÉRREZ-MARCO, J.C. & GARCÍA-BELLIDO, D.C. 2014. A remarkable illaenid trilobite from the Middle Ordovician of Morocco. Bulletin of Geosciences 89(2), 365-374.View article

RAYMOND, P.E. 1916. New and old Silurian trilobites from Southeastern Wisconsin, with notes on the genera of the Illaenidae. Bulletin of the Museum of Comparative Zoology 60(1), 1-41.

RAYMOND, P.E. 1925. Some trilobites of the lower Middle Ordovician of eastern North America. Bulletin of the Museum of Comparative Zoology at Harvard College 67, 1-180.

ROZOVA, A.V. 1960. Upper Cambrian trilobites of Salair. Akademyia nauk SSSR, Sibirskoe otdelenie, Trudy Instituta geologii i geofiziki 5, 1-115. [in Russian]

ROZOVA, A.V. 1968. Biostratigraphic zonation and trilobites of the Upper Cambrian and Lower Ordovician of the north-western Siberian Platform. Akademyia nauk SSSR, Sibirskoe otdelenie, Trudy Instituta geologii i geofiziki 36, 1-196. [in Russian, English translation 1984, by American Publishing Co., New Delhi]

ROZOVA, A.V. 1977. Some Upper Cambrian and Lower Ordovician trilobites from the basins of the Rybnaya, Khantayka, Kureyka and Letnyaya river basins, 54-84. In ZHURAVLEVA, I.T. & ROZOVA, A.V. (eds) Biostratigraphy and fauna of the Upper Cambrian and the boundary strata (New data from the Asiatic part of the U.S.S.R.). Akademyia nauk SSSR, Sibirskoe otdelenie, Trudy Instituta geologii i geofiziki 313. [in Russian]

RŮŽIČKA, R. 1926. Fauna vrstev Eulomových rudního ložiska u Holoubkova (v Ouzkém). Část I. Trilobiti. Rozpravy České akademie věd a umění 35(39), 1-26.

SALTER, J.W. 1867. A monograph of the British trilobites from the Cambrian, Silurian and Devonian formations. Part 4. Palaeontographial Society Monograph 20(86), 177-214.View article

SDZUY, K. 1955. Die Fauna der Leimitz Schiefer (Tremadoc). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 492, 1-74.

SDZUY, K., HAMMANN, W. & VILLAS, E. 2001. The Upper Tremadoc fauna from Vogtendorf and the Bavarian Ordovician of the Frankenwald (Germany). Senckenbergiana lethaea 81(1), 207-261.View article

SHERGOLD, J.H. & SZDUY, K. 1984. Cambrian and early Tremadocian trilobites from Sultan Dag, central Turkey. Senckenbergiana lethaea 65(2/3), 51-135.

SLAVÍČKOVÁ, J. 1999. Family Illaenidae Hawle et Corda, 1847 in the Ordovician of the Prague Basin (Trilobita, Czech Republic). Acta Universitatis Carolinae, Geologica 4(1/2), 361-364.

ŠNAJDR, M. 1957. Klasifikace čeledě Illaenidae (Hawle a Corda) v českém starším paleozoiku. Sborník Ústředního ústavu geologického, Oddíl paleontologický 23, 125-284.

TJERNVIK, T.E. 1956. On the Early Ordovician of Sweden, stratigraphy and fauna. Bulletin of the Geological Institution of the University of Uppsala 36, 107-284.

VANĚK, J. & VALÍČEK, J. 2001. New index of the genera, subgenera, and species of Barrandian trilobites. Part A-B (Cambrian and Ordovician). Palaeontologia Bohemiae 7, 1-49.

VOLBORTH, A. VON 1863. Ueber die mit glatten Rumpfgliedern versehenen russischen Trilobiten, nebst einem Anhange über die Bewegungsorgane und über des Herz derselben. Mémoires de l’Académie Impériale des Sciences de St.-Pétersbourgh, Sér. 7, Tome 6(2), 1-47.

WAHLENBERG, G. 1818. Petrificata Telluris Svecanae. Nova Acta Regiae Societatis Scientarium Uppsaliensis 8, 1-116.

WHITTINGTON, H.B. 1950. Sixteen Ordovician genotype trilobites. Journal of Paleontology 24, 531-565.

WHITTINGTON, H.B. 1963. Middle Ordovician trilobites from Lower Head, western Newfoundland. Bulletin of the Museum of Comparative Zoology at Harvard College 129, 1-118.

WHITTINGTON, H.B. 1965. Trilobites of the Ordovician Table Head Formation, Western Newfoundland. Bulletin of the Museum of Comparative Zoology at Harvard College 132(4), 1-441.

WHITTINGTON, H.B. 1997a. Illaenidae (Trilobita): Morphology of thorax, classification, and mode of life. Journal of Paleontology 71(5), 878-896.

WHITTINGTON, H.B. 1997b. Morphology of the exoskeleton, 1-67. In KAESLER, R.L. (ed.) Treatise on Invertebrate Paleontology. Part O, Revised, Trilobita. Geological Society of America & University of Kansas, Boulder & Lawrence.

WHITTINGTON, H.B. 2000. Stygina, Eobronteus (Ordovician Styginidae, Trilobita): Morphology, classification, and affinities of Illaenidae. Journal of Paleontology 74(5), 879-889.View article

WHITTINGTON, H.B. 2003. The trilobite family Nileidae: Morphology and Classification. Palaeontology 46, 635-646.View article

WHITTINGTON, H.B. & KELLY, S.R.A. 1997. Morphological terms applied to Trilobita, 313-329. In KAESLER, R.L. (ed.) Treatise on Invertebrate Paleontology. Part O, Revised, Trilobita. Geological Society of America & University of Kansas, Boulder & Lawrence.

ZHOU, Z.Y. & FORTEY, R.A. 1986. Ordovician trilobites from north and northeast China. Palaeontographica, Abteilung A 192, 157-210.