Early Devonian polygnathids of Northeast Asia and correlation of Pragian/Emsian strata of the marginal seas of Angarida


Authors: Baranov VV, Slavík L, Blodgett RB

Published in: Bulletin of Geosciences, volume 89, issue 3; pages: 645 - 678; Received 6 September 2013; Accepted in revised form 12 March 2014; Online 9 June 2014

Keywords: Early Devonian, eopolygnathids, Pragian, Emsian, correlation, Northeast Asia, marginal seas, Angarida,

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In late Pragian and early Emsian strata of the Izvestkovyi Karier - II section of Northeast Asia is found a continuous sequence of representatives of the family Polygnathidae which testify to the synthetic nature of the currently used conodont zonal scales. In this time interval are recognized three phylogenetic lineages of polygnathids: Polygnathus sobolevi Po. ivanowskyii, Polygnathus kitabicus Po. bardashevi and Polygnathus settedabanicus Po. nothoperbonus. Fifteen species of polygnathids including mostly newly described taxa are documented here: Polygnathus alkhovikovae sp. nov., Po. arthuri sp. nov., Po. bardashevi sp. nov., Po. dehiscens (Philip & Jackson, 1967) late morphotype, Po. excavatus (Carls & Gandl, 1969), Po. ivanowskyii sp. nov., Po. karsteni sp. nov., Po. kitabicus (Yolkin et al., 1994), Po. lezhoevi sp. nov., Po. michaelmurphyi sp. nov., Po. nothoperbonus (Mawson, 1987), Po. perbonus (Philip, 1966), Po. sobolevi Bardashev, Weddige & Ziegler, 2002, Po. settedabanicus sp. nov., and Po. yakutensis sp. nov. Early Emsian time in Northeast Asia is characterized by an intense speciation and radiation within its conodont fauna during a relatively short interval between the Polygnathus excavatus and Polygnathus nothoperbonus zones. The Pragian Stage (Middle Early Devonian) is characterized by a maximum transgression that led to increased migration of faunal associations of tabulate and rugose corals, brachiopods and conodonts in the marginal seas of Angarida. The position of the lower and upper boundaries of the Pragian Stage in this region is clarified and their correlations are shown with the adjoining Verkhoyansk-Chukotsk (Kolyma region), Taimyr, West Siberian, and Altai-Sayan marginal seas.


Alekseeva, R.E., Gratsianova, R.T., Yolkin, E.A. & Kulkov, N.P. 1970. Stratigrafiya i brakhiopody nizhnego devona severo-vostochnogo Salaira [Stratigraphy and brachiopods of the Lower Devonian of northeastern Salair]. 188 pp. Nauka, ?oskva.

Alekseeva, R.E., Mendbayar, B. & Erlanger, O.A. 1981. Brakhiopody i biostratigrafiya nizhnego devona Mongolii [Brachiopods and stratigraphy of the Lower Devonian of Mongolia]. Trudy sovmestnoi Sovetsko-Mongolskoi paleontologicheskoi ekspedizii 16. 176 pp. Nauka, ?oskva.

Aľkhovik, ?.S. & Baranov, V.V. 2001. Stratigrafiya nizhnego devona severo-vostochnoi Yakutii (Severo-Vostok Rossii) [Stratigraphy of the Lower Devonian of Yakutia (Northeast Russia)]. 149 pp. Yakutskii filial izdatelstva SO RAN, Yakutsk.

Aľkhovik, ?.S. & Ivanowskyi, A.B. 1988. Korally i biostratigrafiya nizhnego devona severo-vostochnoi Yakutii [Corals and biostratigraphy of the Lower Devonian of northeastern Yakutia]. 95 pp. Nauka, ?oskva.

Apekina, L.S. 1984. Conodonty [Conodonts], 73-87. In Kim, A.I, Erina, M.V., Apekina, L.S. & Lessovskaya, A.I. (eds) Biostratigraphiya devona Zeraphano-Gissarskoi gornoi oblasti. Izdatelstvo [Biostratigraphy of Devonian of Zeraphan-Gissarsk Mountains area]. “FAN” Uzbetskoi SSR, Tashkent.

Bakharev, N.K., Izokh, N.G. & Yazikov, A.Y. 2013. New data on the stratigraphy of the Lower and Middle Devonian of Salair (West Siberia, Russia), 25-26. In El Hassani, A., Becker, T. & Tahiri, A. (eds) Abstracts Book, International Field Symposium “The Devonian and Carboniferous of northern Gondwana”, Morocco 2013. Document de l’Institut Scientifique, Rabat 26.

Baranov, V.V. 1991. Konodonty i zonalnaya stratigrafia nizhnego devona Vostochnoi Yakutii [Conodonts and zonation of the Lower Devonian of eastern Yakutia], Regionaľnaya geologiya i poleznye iskopaemye Yakutii], 24-36. Izdatelstvo YaGU, Yakutsk.

Baranov, V.V. 2004. Nizhnii devon Severo-Vostoka Asii (brakhiopody, konodonty, stratigraficheskaya skhema) [The Lower Devonian of Northeast Asia (brachiopods, conodonts, stratigraphic pattern)]. 36 pp. Doctoral (Geol.) Dissertation, IGABM SO RAN, St. Petersburg.

Baranov, V.V. 2009a. Cyklitchnosť sedimentogenesa, etapnosť i stadyinosť razvitia bioty v pozdnem silure i rannem devone Severo-Vostoka Azii [Sedimentogenesis cyclicity, stages, and phases in development of biota in the Late Silurian and Early Devonian of Northeast Asia]. Stratigraphy and Geological Correlation 17(4), 382-395.View article

Baranov, V.V. 2009b. Korrelatsia verkhnesiluriiskikh i nizhnedevonskikh otlozhenii arkticheskikh regionov Azii [Correlation of the Upper Silurian and Lower Devonian beds of the Arctic regions of Asia]. Otechestvennya geologia 5, 75-79.

Baranov, V.V. 2009c. Nizhnaya granitsa devonskoi systemy i granitsy yarusov nizhnego devona na severe Evrazii [The lower boundary of Devonian system and boundaries of stages at north Eurasia]. Nauka i obrazovanie 4, 31-34.

Baranov, V.V. 2012. Zonalnoe raschlenenie nizhnego devona arkticheskikh regionov Evrazii po konodontam [Lower Devonian conodont zonation in Arctic Regions of Eurasia]. Stratigraphy and Geological Correlation 20(2), 68-87. View article

Baranov, V.V. & Blodgett, R.B. 2011. Korrelatsia nizhneprazhskikh otlozhenii (nizhnii devon) Alaski i arkticheskikh regionov Vostochnoi Sibiri [Correlation of the Lower Pragian beds (Lower Devonian) in Alaska and the Arctic regions of Eastern Siberia]. Otechestvennya geologia 5, 38-42.

Baranov, V.V. & Blodgett, R.B. 2012. Problemnye voprosy granits i korrelatsya prazhskogo yarusa (nizhnii devon) okrainnykh morei Angaridy [Problematic questions concerning the boundaries and correlation of the Pragian Stage (Lower Devonian) of the marginal seas of Angarida]. Nauka i obrazovanie 4, 9-13.

Barca, S., Bagnoli, M., Olivieri, R. & Serpagli, E. 1986. New stratigraphic data and evidence of Lower and Upper Devonian based conodonts in Sarrabus area (southeastern Sardinia). Rivista Italiana di Paleontologia e Stratigrafia 92(3), 299-320.

Bardashev, I.A. 1986. Emskie konodonty roda Polygnathus tsentralnogo Tadzhikistana [Emsian conodonts of the genus Polygnathus of Central Tadzhikistan]. Paleontologicheskii Zhurnal 2, 61-66.

Bardashev, I.A. 1991. Devonskaya sistema. Konodonty [The Devonian system. The Conodonts]. Atlas iskopaemoi fauny i flory Tadzhikistana: ordovik, silur, devon, 214-245.

Bardashev, I.A., Weddige, K. & Ziegler, W. 2002. The phylomorphogenesis of some early Devonian platform conodonts. Senckenbergiana lethaea 82, 75-451.View article

Bardashev, I.A. & Ziegler, W. 1992. Conodont biostratigraphy of Devonian deposits of the Shishkat section (Southeastern Tien-Shan, Middle Asian). Courier Forschungsinstitut Senckenberg 154, 1-29.

Barskov, I.S., Kononova, L.I. & Kuzmin, A.V. 1991. Opredeliteľ konodontov devona i nizhnego karbona [Determinant of the conodonts of the Devonian and Carboniferus]. 183 pp. Moskva.

Bassler, R.S. 1925. Classification and stratigraphic use of conodonts. Geological Society of America Bulletin 36, 218-220.View article

Becker, R.T. 2009. Minutes of the SDS Business Meeting, Kitab State Geological Reserve, Uzbekistan. Subcommission on Devonian Stratigraphy Newsletter 24, 12-15.

Becker, R.T. 2012. The uncounted species of Polygnathus - plea for clear tracks through a taxonomic jungle. Subcommission on Devonian Stratigraphy Newsletter 27, 42-48.

Becker, R.T. (ed.) 2013. Subcommission on Devonian Stratigraphy Newsletter 28, 1-76.

Becker, R.T., Gradstein, F.M. & Hammer, O. 2012. The Devonian Period, 559-601. In Gradstein, F.M., Ogg, J.G., Schmitz, M.D. & Ogg, G.M. (eds) The Geologic Time Scale. Elsevier, Oxford.

Boersma, K.T. 1973. Description of certain Lower Devonian platform conodonts of the Spanish Central Pyrenees. Leidse Geologische Mededelingen 49(2), 285-301.

Brock, G.A. & Talent, J.A. 1993. Emsian (Early Devonian) brachiopods from the Ukalunda beds, North Queensland. Memoir of the Association of Australasian Paleontologists 15, 225-248.

Bultynck, P. 1976. Le Silurien supérieur et le Devonien inférieur de la Sierra de Guadarrama (Espagne centrale). Troisieme Partie: Eléments icriodiformes, pelekysgnathiformes et polygnathiformes. Bulletin de L’Institut royal des sciences naturelles de Belgique 49(5), 1-74.

Bultynck, P. 1985. Lower Devonian (Emsian) - Middle Devonian (Eifelian and lowermost Givetian) conodont successions from the Ma‘der and Tafilalt, southern Morocco. Courier Forschungsinstitut Senckenberg 75, 261-256.

Bultynck, P. 1989. Conodonts from the La Grange Limestone (Emsian), Armorican Massif, north-western France. Courier Forschungsinstitut Senckenberg 117, 173-203.

Bultynck, P. & Hollard, H. 1980. Distribution comprée de Conodontes et Goniatites dévoniens des plaines du Dra, du Ma’der et du Tafilalt (Maroc). Aardkundige Mededelingen 1, 1-73.

Carls, P. & Gandl, J. 1969. Stratigraphie und Conodonten des Unter-Devons der östlichen Iberischen Ketten (NE-Spanien). Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 132(2), 144-218.

Carls, P., Slavík, L. & Valenzuela-Ríos, J.I. 2008a. Application to the SDS for revision of the Pragian-Emsian GSSP, 21-22. In Kim, A.I., Salimova, F.A. & Meshchankina, N.A. (eds) International Conference Global Alignments of Lower Devonian Carbonate and Clastic Sequences, SDS and IGCP Project 499 joint field meeting, Contributions. State Committee of the Republic of Uzbekistan on Geology, Kitab State Geological Reserve.

Carls, P., Slavík, L. & Valenzuela-Ríos, J.I. 2008b. Comments on the GSSP for the basal Emsian stage boundary: the need for its redefinition. Bulletin of Geosciences 83(4), 383-390.View article

Carls, P. & Valenzuela-Ríos, J.I. 2000. Faunas, correlations, various boundaries, and opinions concerning the early Emsian sensu lato and a cancellata boundary. Web page of SDS Emsian Working Party. http://www.es.mq.edu.au/mucep/emsian/carls_and_rios/carls_and_rios.htm

Carls, P. & Valenzuela-Ríos, J.I. 2002. Early Emsian Conodonts and associated shelly faunas of the Mariposas Fm (Iberian Chains, Aragon, Spain), 315-333. In García-López, S. & Bastida, F. (eds) Palaeozoic conodonts from northern Spain. Eighth International Conodont Symposium held in Europe, Publicaciones del Instituto Geológico y Minero de Espana, Cuadernos del Museo Geominero, Madrid 1.

Cherkesova, S.V. 1994. Obshaya stratigraficheskaya kharakteristika nizhnedevonskikh otlozhenii tareiskogo razreza [General characteristic of the Lower Devonian beds of the Tareiskay basic section], 7-32. In Cherkesova, S.V., Karatayute-Talimaa, V.N. & Matukhin, R.G. (eds) Stratigrafiya i fauna nizhnedevonskikh otlozhenii tareiskogo opornogo razreza (Taimyr). Nedra, Sankt-Petersburg.

Chlupáč, I., Havlíček, V., Kříž, J., Kukal, Z. & Štorch, P. 1998. Paleozoic of the Barrandian (Cambrian to Devonian). 183 pp. Czech Geological Survey, Praha.

Chlupáč, I., Kříž, J. & Schönlaub, H.P. 1980. Field Trip E. Silurian and Devonian conodonts of the Barrandian, 147-180. In Schönlaub, H.P. (ed.) Second European Conodont Symposium ECOS II. Guidebook - Abstracts. Abhandlungen den Geologischen Bundesanstalt 35.

Chlupáč, I. & Oliver, W.A. 1989. Decision on the Lochkovian - Pragian boundary stratotype (Lower Devonian). Episodes 12(2), 109-113.

Corradini, C. & Corriga, M.G. 2012. A Přídolí-Lochkovian conodont zonation in Sardinia and the Carnic Alps: implications for a global zonation scheme. Bulletin of Geosciences 87(4), 635-650.View article

Drygnat, D. & Szaniawski, H. 2012. Lochkovian conodonts from Podolia, Ukraine and their stratigraphic significance. Acta Palaeontologica Polonica 57(4), 833-861.View article

Dubatolov, V.N., Krasnov, V.I., Rotanov, L.S., Bogush, O.I., Isaev, G.D., Serdyuk, Z.Y. & Stepanov, S.?. 1990. Litostratigraficheskoe rastchlenenie i korrelatsia razrezov parametricheskikh i razvedochnykh skvazhin [Litostratigraphical zonation and correlation of the parametrically and prospect holes], 5-37. In Tesakov, Y.I. (ed.) Stratigrafiya paleozoiskikh otlozhenii yugo-vostoka Zapadno-Sibirskoi plity. Nauka, Novosibirsk.

Egorova, L.I. & Savitskyi, B.E. 1969. Stratigrafia i biofatsii kembria Sibirskoi platformy (zapadnoe Prianabarye) [Stratigraphy and biofacies of the Cambrian of the Siberian Platform (western Prianabarye)]. Trudy NIIGGIMS, Seria stratigrafia i paleontologia 43. 408 pp. Nedra, Moskva.

Erina, M.V. 2007. Conodonts. In Kim, A.I., Salimova, F.A. & Meshchankina, N.A. (eds) Paleontological Atlas of Phanerozoic faunas and ?ora of Uzbekistan. Vol. 1. Paleozoic (Cambrian, Ordovician, Silurian, Devonian, Carboniferous, Permian). 1473 pp. IMR, Tashkent.

Fahraeus, L.E. 1971. Lower Devonian conodonts from the Michelle and Prongs Creek Formations Yukon Territory. Journal of Paleontology 45, 665-683.

Gagiev, M.K. 1987. Stratigrafiya i konodonty nizhnedevonskikh otlozhenii Severo-Vostoka SSSR [Stratigraphy and conodonts of the lower Devonian beds of Northeast USSR], SVKNII DVNTS AN SSSR. 40 pp. Preprint, Magadan.

Gagiev, M.K. 1995. Stratigrafiya devona i nizhnego karbona Omulevskogo podnyatia (Severo-Vostok Azii) [Stratigraphy of the Devonian and Lower Carboniferous of Omulevskay Mountains (Northeast Asia)]. 196 pp. SVKNII DVO RAN, Magadan.

Gagiev, M.K., Dubatolov, V.N. & Terekh, G.A. 1990. Opornyi razrez nizhnego devona Prokolymskogo podnyatia [Basic section of the Lower Devonian of Kolyma Uplift]. 24 pp. SVKNII DVO SSSR, Preprint, Magadan.

Gagiev, M.K., Rodygin, S.A. & Timofeeva, O.B. 1987. Zonaľnoe raschlenenie i korrelatsia nizhne-srednedevonskikh otlozhenii Salaira i Severo-Vostoka SSSR po konodontam [Zonation and correlation of the Lower and Middle Devonian deposits of Salair and Northeast USSR]. 55 pp. RIO DVNTs, Preprint, Magadan.

García-López, S. & Alonso-Menendez, C. 1994. Conodontos del Grupo La Vid limites Praguinse / Emsiense y Emsiense Inferior/Emsiense Superior, Zona Cantabrica (NO de Espana). Revista Espanola de Micropaleontologia 26(2), 81-97.

Gratsianova, R.T. 1967. Brakhiopody i stratigraphy nizhnego devona Gornogo Altaya [Brachiopods and stratigraphy of the Lower Devonian of Gornyi Altai]. 177 pp. Nauka, ?oskva.

Hinde, G.L. 1879. On conodonts from the Chazy and Cincinati group of the Cambro-Silurian, and from the Hamilton and Genesee Shale divisions of the Devonian, in Canada and the United States. Quarterly Journal of the Geological Society of London 35(3), 351-369.View article

Hladil, J., Čejchan, P., Gabašová, A., Táborský, Z. & Hladíková, J. 1996. Sedimentology and orientation of tentaculite shells in turbidite lime mudstone to packstone: Lower Devonian, Barrandian, Bohemia. Journal of Sedimentary Research 66, 888-899.

Hladil, J. & Kalvoda, J. 1997. A short range anomaly in the earliest Emsian sedimentation of the Barrandian: possible reflection of widely controlled or global event. Subcommission on Devonian Stratigraphy, Newsletter 13, 37-38.

Hladil, J., Slavík, L., Vondra, M., Koptíková, L., Čejchan, P., Schnabl, P., Adamovič, J., Vacek, F., Vich, R., Lisá, L. & Lisý, P. 2011. Pragian-Emsian successions, Uzbekistan and Bohemia: magnetic susceptibility logs and dynamic time alignment. Stratigraphy 8(4), 217-235.

Holcová, K. & Slavík, L. 2013. The morphogroups of small agglutinated foraminifera from the Devonian carbonate complex of the Prague Synform, (Barrandian area, Czech Republic). Palaeogeography, Palaeoclimatology, Palaeoecology 386, 210-224.View article

Izokh, N.G. 1998. Konodonty i zonalnoe raschlenenie emsskikh otlozhenii Salaira [Conodonts and zonal division of the Emsian division of the Emsian]. 26 pp. Cand. Sci. (Geol.) dissertation. IGNiG SO RAN. Novosibirsk.

Izokh, N.G., Yolkin, E.A., Weddige, K., Erina, M.V. & Valenzuela-Ríos, J.I. 2011. Late Pragian and Early Emsian conodont polygnathid species from the Kitab state geological reserve sequences (Zeravshan-Gissar Mountainous Area, Uzbekistan). Novosti geologii i stratigrafii 15 (Prilozhenie k zhurnalu “Geologia i geofizika”) 52, 49-63.

Jansen, U. 2012. On the traditional Siegenian-Lower Emsian successions in the Rheinish Slate Mountains, in special consideration of the basal Emsian boundary and its supraregional correlation. Subcommission on Devonian Stratigraphy Newsletter 27, 21-27.

Jansen, U., Becker, G., Plodowski, G., Schindler, E., Vogel, O. & Weddige, K. 2004. Pragian and Emsian near Aouinet Torkoz (SW Dra Valley, Morocco). Devonian of the western Anti Atlas: correlations and events. Document de l’Institut Scientifique, Rabat 19, 75-84.

Klapper, G. 1969. Lower Devonian conodonts sequence, Royal Creek, Yukon Territory, and Devon Island, Canada; with a section on Devon Island stratigraphy by A.R. Ormiston. Journal of Paleontology 43, 1-27.

Klapper, G. 1977. Lower and Middle Devonian conodont sequence in central Nevada; with contributions by Johnson, D.B. University of California, Riverside Campus Museum Contributions 4, 33-54.

Klapper, G. & Johnson, D.B. 1975. Sequence in conodont genus Polygnathus in Lower Devonian at Lone Mountain, Nevada. Geologica et Palaeontologica 9, 65-83.

Klapper, G. & Johnson, D.B. 1980. Endemism and dispersal of Devonian conodonts. Journal of Paleontology 52, 400-455.

Klapper, G. & Philip, G.M. 1971. Devonian conodont apparatuses and their vicarious skeletal elements. Lethaia 4, 129-452.View article

Klapper, G. & Vodrážková, S. 2013. Ontogenetic and intraspecific variation in the late Emsian - Eifelian (Devonian) conodonts Polygnathus serotinus and P. bultyncki in the Prague Basin (Czech Republic) and Nevada (western U.S.). Acta Geologica Polonica 63(2), 153-174.View article

Klapper, G., Ziegler, W. & Mashkova, T.V. 1978. Conodonts and correlation of Lower-Middle Devonian boundary beds in the Barrandian area of Czechoslovakia. Geologica et Palaeontologica 12, 103-116.

Klets, A.G. 2005. Verkhnii paleozoi okrainnykh morei Angaridy [Upper Paleozoic of the marginal seas of Angarida]. 240 pp. Geo, Novosibirsk.

Koptíková, L., Bábek, O., Hladil, J., Kalvoda, J. & Slavík, L. 2010b. Stratigraphic significance and resolution of spectral reflectance logs in Lower Devonian carbonates of the Barrandian area, Czech Republic; a correlation with magnetic susceptibility and gamma-ray logs. Sedimentary Geology 225, 83-98.View article

Koptíková, L., Hladil, J., Slavík, L., Bábek, O. & Čejchan, P. 2010a. Fine-grained non-carbonate particulates embedded in neritic to pelagic limestones (Lochkovian to Emsian, Prague Synform, Czech Republic): Composition, provenance and links to magnetic susceptibility and gamma-ray logs. Geologica Belgica 13(4), 407-430.

Koren’, T.N. 2006. Zonaľnaya stratigrafiya fanerozoya Rossii [The Phanerozoic Zonal Stratigraphy of Russia]. 256 pp. VSEGEI, St. Petersburg.

Krylova, A.K. 1962. Stratigrafiya i brakhiopody devonskoi sistemy Sibirskoi platformy [Stratigraphy and brachiopods of the Devonian system of Siberian Platform]. 108 pp. Gostoptekhizdat, Leningrad.

Kulkov, N.P. 1963. Brakhiopody solovikhinskikh sloev nizhnego devona Gornogo Altaya [Brachiopods of the Solovyikhinsky layers of Lower Devonian of Gornyi Altai]. 131 pp. Nauka, ?oskva.

Kulkov, N.P. & Peregoedov, L.G. 1990. Klass Articula. Stratigrafiya paleozoiskikh otlozhenii yugo-vostoka Zapadno-Sibirskoi plity [Class Articulata. Stratigraphy of the Paleozoic beds of South-East West-Siberian plateau], 70-106. In Tesakov, Y.I. (ed.) Stratigrafiya paleozoiskikh otlozhenii yugo-vostoka Zapadno-Sibirskoi plity. Nauka, Novosibirsk.

Lane, H. R. & Ormiston, A. R. 1979. Siluro-Devonian biostratigraphy of the Salmontrout River area, east-central Alaska. Geologica et Palaeontologica 13, 39-96.

Ladeux, H. & Morzadec, P. with. contr. of Bultynck, P. & Walliser, O.H. 1979. Massiv Armorican. SDS Field Excursion Guide, 1-27.

Martínez-Pérez, C. 2010. Icriodus riosi, nueva especie de conodonto en el Devónico Inferior del Pirineo Central espanol [Icriodus riosi, a new conodont species from the Lower Devonian of the Spanish Central Pyrenees]. Revista Espanola de Paleontología 25(2), 139-147.

Martínez-Pérez, C. & Valenzuela-Ríos, J.I. 2005. Conodontos del Límite Praguiense/Emsiense (Devónico Inferior) en la sección Isábena 1 (Huesca, Pirineo Aragonés), 287-319. In Menendéz, G, Martínez-Pérez, C, Ros, S., Botella, H. & Plasencia, P. (eds) Miscelánea Paleontológica. Publicaciones del Seminario de Paleontología de Zaragoza (SEPAZ), Zaragoza 6.

Martínez-Pérez, C., Valenzuela-Ríos, J.I. & Botella, H. 2010. Polygnathus rosae n. sp. (Conodonta) and its biostratigraphical correlation potential (lower Emsian, Lower Devonian) in the Spanish Central Pyrenees. Rivista Italiana di Paleontologia e Stratigrafia 116(3), 273-281.

Martínez-Pérez, C., Valenzuela-Ríos, J.I., Navas-Parejo, P., Liao, J.-C. & Botella, H. 2011. Emsian (Lower Devonian) Polygnathids (Conodont) succession in the Spanish Central Pyrenees. Journal of Iberian Geology 37(1), 45-64.View article

Mashkova, T.V. & Apekina, L.S. 1978. Conodonts, pls 73-78. In Kim, A.I., Yolkin, E.A., Erina, M.V. & Gratsianova, R.T. (eds) Field Session of the International Subcommission on Devonian Stratigraphy, Samarkand, USSR. Type sections of the Lower and Middle Devonian boundary beds in Middle Asia, together with A Guide to Field Excursions. Atlas of palaeontological plates. Supplement to Guide of the Field Excursion. Tashkent.

Mashkova, T.V. & Apekina, L.S. 1980. Prazhskie polignatusy (konodonty) zony dehiscens Sredney Asii [Pragian polygnathids (Conodonts) of dehiscens Zone of Middle Asia]. Paleontologicheskii Zhurnal 3, 135-140.

Mashkova, T.V. & Apekina, L.S. 1981. Drevneishie polignatusy Zeravshanskogo khrebta - pokazateli granitsi nizhnego i srednego devona [The oldest Polygnathuses from the Zeravshan Range as the Lower-Middle Devonian boundary indicators]. Izvestia Akademii nauk SSSR, Seria geologicheskaya 2, 88-94.

Mashkova, T.V. & Sobolev, N.N. 1980. Drevneishie polignatusy Novoi Zemli - pokazateli granitsy nizhnego-srednego devona [The oldest polygnathuses of Novaya Zemlya as a parameters of the boundary of the Lower and Middle Devonian]. Izvestia Akademii nauk SSSR, Seria geologicheskaya 5, 56-61.

Mastandrea, A. 1984. Biostratigraphic remarks on Early Devonian conodonts from Corti Baccas III section (NW Sardinia). Bollettino della Societa Paleontologica Italiana 23(2), 259-267.

Matukhin, R.G., Menner, V.V., Sokolov, P.N. & Reshetnyak, D.R. 1995. Stratigraficheskaya osnova devonskoi sistemy Sibirskoi platformy [Stratigraphical base of the Devonian system of Siberian Platform], 39-96. SNIIGGIMS, Novosibirsk.

Mavrinskaya, T. & Slavík, L. 2013. Correlation of Early Devonian (Lochkovian-early Pragian) conodont faunas of the South Urals (Russia). Bulletin of Geosciences 88(2), 283-296.View article

Mawson, R. 1995. Early Devonian polygnathid conodont lineages with special reference to Australia. Courier Forschungsinstitut Senckenberg 182, 389-398.

Mawson, R. 1987. Documentation of conodont assemblages across the Early Devonian-Middle Devonian boundary, Broken River Formation, North Queensland, Australia. Courier Forschungsinstitut Senckenberg 92, 251-273.

Mawson, R. 1998. Thoughts on Late Pragian-Emsian polygnathid evolution: documentation and discussion, 201-208. In Szaniawski, H. (ed.) Proceedings of the Sixth European Conodont Symposium (ECOS VI). Palaeontologia Polonica 58.

Mawson, R. & Talent, J.A. 2003. Conodont faunas from sequences on or marginal to the Anakie Inlier (Central Queensland, Australia) in relation to Devonian transgressions. Bulletin of Geosciences 78(4), 335-358.

Mawson, R., Talent, J.A., Brock, G.A. & Engelbretsen, M.J. 1992. Conodont data in relation to sequences about the Pragian-Emsian Boundary (Early Devonian) in south-eastern Australia. Proceedings of the Royal Society of Victoria 104(1), 23-56.

McGregor, D.C. & Uyeno, T.C. 1972. Devonian spores and conodonts of Melville and Bathurst Islands, District of Franklin. Geological Survey of Canada, Paper 71-13, 1-37.

Menner, V.V., Nikolaev, A.A. & Rzonsnitskaya, M.A. 1973. Severo-Vostok SSSR [Northeast USSR], 157-184. In Nalivkin, D.V., Rzonsnitskaya, M.A. & Markovskyi, B.P. (eds) Devonskaya sistema. Kniga 2. Nedra, Moskva.

Mironova, N.V. 1974. Rannedevonskie tabulaty Gornogo Altaya i Salaira [Early Devonian tabulates of Gornyi Altai and Salair]. V. 168. 164 pp. Zapadno-Sibirskoe knizhnoe izdatelstvo, SNIIGGIMS, Novosibirsk.

Murphy, M.A. 1989. Lower Pragian boundary (Lower Devonian) and its application in Nevada. Courier Forschungsinstitut Senkenberg 117, 61-70.

Murphy, M.A. 2005. Pragian conodont zonal classification in Nevada, Western North America. Revista Espanola de Paleontología 20, 177-206.

Murphy, M.A. & Matti, J.C. 1982. Lower Devonian conodonts (hesperius-kindlei zones), central Nevada. University of California Publications in Geological Sciences 123, 1-83.

Murphy, M.A. & Valenzuela-Ríos, J.I. 1999. Lanea new genus, lineage of Early Devonian conodonts. Bolletino della Societa Paleontologica Italiana 37, 321-334.

Nikiforova, O.I. 1960. Novyi vid gyurikhellid Arktiki [New species of gurichellids of Arctica], 399-402. In Markovskyi, B.P. (ed.) Novye vidy drevnikh rastenii i bespozvonochnykh SSSR. Chasť 1. Gosgeoltekhizdat, Moskva.

Nikiforova, O.I. & Andreeva, O.N. 1961. Stratigrafiya ordovika i silura Sibirskoi platformy i eyo paleontologicheskoe obosnovanie [Stratigraphy Ordovician and Silurian of the Siberian Platform and it paleontological substantiation]. 142 pp. Gostoptekhizdat, Leningrad.

Olivieri, R. & Serpagli, E. 1990. Latest Silurian-early Devonian conodonts from the Mason Porcus Section near Fluminimaggiore, Southwestern Sardinia. Bollettino della Societa Paleontologica Italiana 29, 59-76.

Pepelyaev, B.V. & Simakov, K.V. 1974. K stratigrafii devonskikh otlozhenii Prikolymskogo podnyatia [To stratigraphy of the Devonian rocks of Kolyma reising], 96-102. In Nikolaev, A.A. & Simakov, K.V. (eds) Dokembrii i paleozoi Severo-Vostoka SSSR, Tezisy dokladov. Magadan.

Perry, D.G., Klapper, G. & Lenz, A.C. 1974. Age of the Ogilvie formation (Devonian), northern Yukon: based primarily on the occurrence of brachiopods and conodonts. Canadian Journal of Earth Sciences 11, 1055-1097.View article

Philip, G.M. 1966. Lower Devonian conodonts from the Buchan Group, eastern Victoria. Micropaleontology 12(4), 441-460.View article

Philip, G.M. & Jackson, J.H. 1967. Lower Devonian subspecies of the conodont Polygnathus linguiformis Hinde from southeastern Australia. Journal of Paleontology 41(5), 1262-1266.

Rzhonsnitskaya, M.A. 1968. Biostratigraphia devona okrain Kuznetskogo basseina. Stratigrafiya [Biostratigraphy of Devonian of the margin Kuznetsk basin]. T. 1. 286 pp. Nedra, Leningrad.

Savage, N.M. 1977a. Lower Devonian conodonts from the Karheen Formation, southeastern Alaska. Canadian Journal of Earth Sciences 14, 278-284.View article

Savage, N.M. 1977b. Lower Devonian conodonts from the Gazelle Formation, Klamath Mountains, Northern California. Journal of Paleontology 51(1), 57-62.

Savage, N.M. 1981. Early Devonian conodonts from Kasaan Island, southeastern Alaska. Journal of Paleontology 55, 848-853.

Savage, N.M., Blodgett, R.B. & Jaeger, H. 1985. Conodonts and associated graptolites from the Lower Devonian of east-central Alaska and western Yukon Territory. Canadian Journal of Earth Sciences 22(12), 1880-1883.View article

Savage, N.M., Churkin, M. Jr. & Eberlein, G.D. 1977. Lower Devonian conodonts from Port St. Nicholas, southeastern Alaska. Canadian Journal of Earth Sciences 14, 2928-2936.View article

Schönlaub, H.P. 1985. Devonian conodonts from section Oberbuchach II in the Carnic Alps. Courier Forschungsinstitut Senckenberg 75, 363-374.

Sharkova, T.T. 1981. Siluriiskie i devonskie tabulata Mongolii [Silurian and Devonian tabulats of Mongolia]. Trudy sovmestnoi Sovetsko-Mongolskoi paleontologicheskoi ekspeditsii 14. 102 pp. Nauka, Moskva.

sine 1982. Reshenia Vsesoyuznogo soveschania po razrabotke unifitscirovannykh stratigraficheskikh schem dokembria, paleozoya i chetvertichnoi sistemy Srednei Sibiri, chast’ 2 (srednii i verkhnii paleozoi) [Solutions-Union meeting on development of unified stratigraphic schemes Precambrian, Paleozoic and Quaternary systems of Central Siberia]. 129 pp. Novosibirsk.

sine 2012. Regionalnaya stratigraficheskaya schema devonskikh obrazovanii vostochnoi chasti Altae-Sayanskoi oblasti [Regional stratigraphic scheme of the eastern part of Altai-Sayan area]. 52 pp. SNIIGGiMS, Novosibirsk.

Slavík, L. 2004a. The Pragian-Emsian conodont successions of the Barrandian area: search of an alternative to the GSSP polygnathids-based correlation concept. Geobios 37(4), 454-470.View article

Slavík, L. 2004b. A new conodont zonation of the Pragian in the stratotype area (Barrandian, central Bohemia). Newsletters on Stratigraphy 40, 39-71.View article

Slavík, L., Carls, P., Hladil, J. & Koptíková, L. 2012. Subdivision of the Lochkovian Stage based on conodont faunas from the stratotype area (Prague Synform, Czech Republic). Geological Journal 47, 616-631.View article

Slavík, L. & Hladil, J. 2004. Lochkovian/Pragian GSSP revisited: evidence about conodont taxa and their stratigraphic distribution. Newsletters on Stratigraphy 40(3), 137-153.View article

Slavík, L., Hladil, J., Valenzuela-Ríos, J.I., Carls, P., Koptíková, L. & Čejchan, P. 2011. Prospect for subdivision of the Pragian Stage based on data from the type area, 144-145. In Obut, O.T. & Kiprianova, T.P. (eds) Biostratigraphy, paleogeography and events in Devonian and Lower Carboniferous (SDS/IGCP 596 joint field meeting): Contributions of the International Conference in memory of Evgeny A. Yolkin. Publishing house of SB RAS, Ufa, Novosibirsk.

Slavík, L., Valenzuela-Ríos, J.I., Hladil, J. & Carls, P. 2007. Early Pragian conodont-based correlations between the Barrandian area and the Spanish Central Pyrenees. Geological Journal 42, 499-512.View article

Sobolev, N.N. 1984. Konodonty iz nizhne-srednedevonskikh otlozhenii Novoi Zemli [Conodonts from the Lower and Middle Devonian of the Novaya Zemlya], 58-85. In Novaya Zemlya na rannikh etapakh geologicheskogo razvitia. PGO “Sevmorgeologia”, Leningrad.

Sobolev, N.N. 1994. Konodonty [Conodonts], 93-99. In Cherkesova, S.V., Karatayute-Talimaa, V.N. & Matukhin, R.G. (eds) Stratigrafiya i fauna nizhnedevonskikh otlozhenii tareiskogo opornogo razreza (Taimyr). Nedra, Sankt-Petersburg.

Sobolev, N.N. & Evdokimova, I.O. 2013. Obtchaya stratigraficheskaya shkala devonskoi systemy: sostoyanie i problemy [General stratigraphic scale of the Devonian system: current state and problems], 130-148. In Fedonkin, M.A. (ed.) Obshaya stratigraficheskaya shkala Rossii: sostoyanie i perspectivy obustroistva. Vserossiiskaya konferencia. Moskva, 23-25 maya, 2013 g. Sbonik statei. GIN RAN, Moskva.

Sweet, W.?. 1988. The Conodonta: morphology, taxonomy, paleoecology and evolutionary history of a long-extinct animal phylum. Oxford Monographs on Geology and Geophysics 10. 212 pp. Clarendo Press, Oxford.

Talent, J.A. & Mawson, R. 1999. North-Eastern Molong Arch and Adjacent Hill End Trough (Eastern Australia). Mid Palaeozoic conodont date and implications. North Gondwana Stratigraphy and Biota 54, 49-105.

Talent, J.A., Mawson, R., Aitchison, J.C., Becker, R.T., Bell, K.N., Bradshaw, M.A., Burrow, C.J., Cook, A.G., Dargan, G.M., Douglas, J.G., Edgecombe, G.D., Feist, R.M., Jones, P.J., Long, J.A., Phillips-Ross, J.R., Pickett, J.W., Playford, G., Rickards, R.B., Webby, B.D., Winchester-Seeto, T., Wright, A.J., Young, G.C. & Zhen, Y.-Y. 2000. Devonian palaeobiogeography of Australia and adjoining regions. Memoirs of the Association of Australasian Palaeontologists 23, 167-257.

Tarabukin, V.P. 1989. Konodonty i biostratigrafia pogranichnykh oilozhenii Silura i Devona Selennyakhskogo krayzha [Conodonts and biostratigraphy of the Silurian and Devonian boundary deposits of Selennyakh Range], 111-120. Izdatelstvo YaGU, Yakutsk.

Tarabukin, V.P. 1992. Biostratigrafia i konodonty devonskikh otlozhenii Vostchnoi Yakutii [Biostratigraphy and conodonts of the Devonian deposits of eastern Yakutia]. 21 pp. Cand. Sci. (Geol.) Dissertation, Moskva.

Timofeeva, O.B. 1973. O nakhodke konodontov v nizhnem devone Kuznetskogo basseina [About a find conodonts in the Lower Devonian of Kuznetsk basin], 236-239. In Nalivkin D.V. (ed.) Stratigrafiya nizhnego i srednego devona. Tom 2. Nauka, Leningrad.

Uyeno, T.T. 1990. Biostratigraphy and conodont faunas of Upper Ordovician through Middle Devonian rocks, Eastern Arctic Archipelago. Bulletin of Geological Survey of Canada, Paper 401, 1-211.

Uyeno, T.T. & Klapper, G. 1980. Summary conodont biostratigraphy of the Blue Fiord and Bird Fiord Formations (Lower Middle Devonian) at the type and adjacend areas, southwestern Ellesmere Island, Canadian Arctic Archipelago; in Current Research. Part C. Geological Survey of Canada, Paper 80-1C, 81-93.

Valenzuela-Ríos, J.I. & Carls, P. 2010. Brief comments on the Future Pragian Subdivision and Revision of the Emsian base. Subcommission on Devonian Stratigraphy Newsletter 25, 19.

Walliser, O.H. & Kim, A.I. 2001. On lowermost Emsian tentaculites from the Zinzilban boundary section and their utility for correlation. Document submitted to the Subcommission on Devonian Stratigraphy, May 2001. 2 pp. Frankfurt am Main.

Wang, C-Y. 1981. Conodonts from Devonian Ertang Formation in central Guangxi. Acta Palaeontologica Sinica 20(5), 400-405.

Wang, C-Y. & Wang, Z.H. 1981. Conodont sequence in China (Cambrian-Triassic). 12th Annual Conference of the Palaeontological Society of China, Selected Papers, 105-115.

Wang, C.-Y. & Ziegler, W. 1983. Conodonten aus Tibet. Neues Jahrbuch für Geologie und Paläontolgie, Monatshefte 1983(2), 65-79.

Weddige, K. 1977. Die Conodonten der Eifel-Stufe im Typusgebiet und in benachbarten Faziesgebieten. Senckenbergiana lethaea 58, 271-419.

Weddige, K. 1987. The Lower Pragian boundary (Lower Devonian) based on the conodont species Eognathodus sulcatus. Senckenbergiana lethaea 67, 479-487.

Weddige, K. & Ziegler, W. 1977. Correlation of Lower/Middle Devonian boundary beds. Newsletters on Stratigraphy 6(2), 67-84.View article

Weddige, K. & Ziegler, W. 1979. Evolutionary patterns in Middle conodont genera Polygnathus and Icriodus. Geologica et Paleontologica 13(3), 157-164.

Weyant, M. 1992. Decouverte de deux conodonts de zone du Praguien (Devonien inférieur) dans le Massif Armoricain. Geobios 25(2), 279-282.View article

Xiong, J.-F. 1983. Devonian conodonts, 301-320. In Chengdu Institute of Geology and Mineral Resources (ed.) Paleontological Atlas of Southwest China. Volume of microfossils. Geological Publishing House, Beijing.

Yolkin, E.A., Apekina, L.S., Erina, M.V., Izokh, N.G., Kim, A.I., Talent, J.A., Walliser, O.H., Weddige, K., Werner, R. & Ziegler, W. 1989. Polygnathid lineages across the Pragian-Emsian Boundary, Zinzilban Gorge, Zerafshan, USSR. Courier Forschungsinstitut Senckenberg 110, 237-246.

Yolkin, E.A., Bakharev, N.K., Gratsianova, R.T., Zheltonogova, V.A., Izokh, N.G., Yazikov, A.Y., Vjushkova, L.V., Mezentseva, O.P., Petrosyan, N.M. & Timpkhina, I.G. 1987. Stratotypicheskie razrezy nizhnego i srednego devona Salaira (Telengitskii nadgorizont: karbonatnye facii) [Stratotypical sections of the Lower and Middle Devonian of Salair (Telengitskay supergorizon. Carbonate facies]. 194 pp. IGiG SO RAN SSSR, Novosibirsk.

Yolkin, E.A. & Izokh, N.G. 1988. Zonalnye vidy konodontov v tipovykh vykhodakh telengitskogo nadgorizonta (devon, Salair) [The zonal species of conodonts in typical exits of the Telengitskyi Superhorizonte (Devonian, Salair)], 3-17. In Yolkin, E.A. & Kanygin, A.V. (eds) Fauna i stratigraphia paleozoya Srednei Sibiri i Urala. Nauka, Novosibirsk.

Yolkin, E.A., Izokh, N.G., Weddige, K., Erina, M.V., Valenzuela-Ríos, J.I. & Apekina, I.L. 2011. Eognathodid and polygnathid lineages from the Kitab state geological reserve sections (Zeravshan-Gissar Moutainous Area, Uzbekistan) as the bases for improvements of Pragian-Emsian standard conodont zonation. Novosti paleontologii i stratigrafii 15, Prilozhenie k zhurnaly “Geologia i geofizika” 52, 37-45.

Yolkin, E.A., Kim, A.I., Weddige, K., Talent, J.A. & House, M.R. 1997. De?nition of the Pragian/Emsian Stage boundary. Episodes 20(3), 235-240.

Yolkin, E.A., Weddige, K., Izokh N.G. & Erina, M.V. 1994. New Emsian conodont zonation (Lower Devonian). Courier Forschungsinstitut Senckenberg 168, 139-157.

Ziegler, W. (ed.) 1977. Catalogue of Conodonts 3. 570 pp. E. Schweizerbart’sche Verlagsbuchhandlung, Stuttgart.

Ziegler, W. & Wang, C.-Y. 1985. Sihongshan section, a regional reference section for the Lower-Middle and Middle-Upper Devonian boundaries in East Asia. Courier Forschungsinstitut Senckenberg 75, 17-37.

Ziegler, W. & Weddige, K. 1999. Zur Biologie, Taxonomie und Chronologie der Conodonten. Paläontologische Zeitschrift 73, 1-38.View article