Graptolites of the Králův Dvůr Formation (mid Katian to earliest Hirnantian, Czech Republic)

 

Authors: Kraft P, Štorch P, Mitchell CE

Published in: Bulletin of Geosciences, volume 90, issue 1; pages: 195 - 225; Received 27 March 2013; Accepted in revised form 5 November 2014; Online 19 January 2015

Keywords: graptolite, Ordovician, Katian, Hirnantian, Prague Basin, biostratigraphy,

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Abstract

Ten graptolite species have been recognized by a general revision of all the material (both historical and new) collected from the Králův Dvůr Formation of the Prague Basin. One species belongs to the dicranograpid genus Dicellograptus and the remaining are biserial taxa assigned to the diplograptid genera Diplograptus and Anticostia, the lasiograptids Paraplegmatograptus and Phormograptus, the climacograptid Styracograptus, and the neodiplograptid genera Normalograptus and Metabolograptus. Although graptolite specimens are rare in these strata, there does appear to be a regular succession among the species, which results in a relatively low diversity of graptolites throughout the formation. The maximum species diversity is in the upper part preceding the distinct carbonate Perník Bed sedimentation. Three of the four species that occur in the lower part of the formation are endemic to the study region, however, Anticostia teres occurs in P. linearis Zone strata in Baltica (Bornholm and Sweden). Three of the four species that occur in the middle and upper part of the unit are more widely distributed species that occur also in the late Katian of the paleotropics. The uppermost Králův Dvůr Formation is marked by impoverishment that reduced the fauna to a single cosmopolitan species that occupies a narrow interval below the first early Hirnantian diamictite. This graptolite distribution pattern appears to reflect changing global environmental conditions, including a late Katian climate amelioration (with expanded geographic ranges of several paleotropical graptolite species) followed by cooling and climate deterioration during latest Katian and early Hirnantian interval. Based on the new data, a biostratigraphic scheme has been modified and the Králův Dvůr Formation is inferred to be of mid Katian (P. linearis) to the earliest Hirnantian (early M. extraordinarius) age.

References

Ainsaar, L., Kaljo, D., Martma, T., Meidla, T., Männik, P., Nolvak, J. & Tinn, O. 2010. Middle and Upper Ordovician carbon isotope chemostratigraphy in Baltoscandia: A correlation standard and clues to environmental history. Palaeogeography, Palaeoclimatology, Palaeoecology 294(3-4), 189-201.View article

Apollonov, M.K., Bandaletov, S.M. & Nikitin, J.F. 1980. The Ordovician-Silurian Boundary in Kazakhstan. 232 pp. Nauka of Kazakh SSR, Alma-Ata. [in Russian]

Armstrong, H.A., Baldini, J., Challands, T.J., Gröcke, D.R. & Owen, A.W. 2009. Response of the Inter-tropical Convergence Zone to Southern Hemisphere cooling during Upper Ordovician glaciation. Palaeogeography, Palaeoclimatology, Palaeoecology 284(3-4), 227-236.View article

Barrande, J. 1870. Défense des Colonies. 4. 186 pp. Prague, Paris.

Bergström, S.M., Chen Xu, Gutiérrez-Marco, J.C. & Dronov, A. 2009. The new chronostratigraphic classification of the Ordovician System and its relations to major regional series and stages and to ?13C chemostratigraphy. Lethaia 42(1), 97-107.View article

Bouček, B. 1928. O vrstvách zahořanských - d???????????????????? českého ordoviku. Rozpravy České akademie věd a umění, Třída II (matematicko-přírodovědecká) 37(33), 1-59. [in Czech; concurrent French version]

Brenchley, P.J. & Štorch, P. 1989. Environmental changes in the Hirnantian (upper Ordovician) of the Prague Basin, Czechoslovakia. Geological Journal 24(3), 165-181.View article

Brussa, E.D., Toro, B.A., Mitchell, C.E. & Astini, R.A. 2003. Biostratigraphic remarks on the Late Ordovician graptolite fauna from the area around Estancia Canota, 53-57. In Albanesi, G.L., Beresi, M.S. & Peralta, S.H. (eds) Ordovician from the Andes. INSUGEO, Serie Correlación Geológica 17. 549 pp. Consejo Nacional de Investigaciones Científicas y Técnicas, Facultad de Ciencias Naturalles e Instituto Miguel Lillo, Universidad Nacional de Tucumán, Tucuman

Bruvo, M. 2005. Graptolitzoneringen i Dicellograptus Skifern (Ovre Ordovicium) pa Bornholm, Danmark. 105 pp. MSc. thesis, University of Copenhagen.

Budil, P., Fatka, O., Kolář, P. & David, M. 2011a. Arthrorhachis Hawle & Corda, 1847 (Agnostida) in the Prague Basin (Barrandian area, Czech Republic) revisited. Bulletin of Geosciences 86(4), 707-724.View article

Budil, P., Fatka, O., Mergl, M. & David, M. 2011b. Trilobite biostratigraphy of the Králův Dvůr Formation (upper Katian, Prague Basin, Czech Republic): global faunal changes or facies-related distribution? Geologické výzkumy na Moravě a ve Slezsku 18(1), 4-10.

Carruthers, W. 1867. Graptolites: their structure and systematic position. Intellectual Observer 11(4), 10(5), 64, 65.

Chen Xu, Fan Jun-xuan, Melchin, M.J. & Mitchell, C.E. 2005. Hirnantian (late Ordovician) graptolites from the Upper Yangtze Region, China. Palaeontology 48, 235-280.View article

Chen Xu, Rong Jia-yu, Fan Jun-xuan, Zhan Ren-bin, Mitchell, C.E., Harper, D.A.T., Melchin, M.J., Peng Ping’an, Finney, S.C. & Wang Xiao-feng 2006. The Global Boundary Stratotype Section and Point (GSSP) for the base of the Hirnantian Stage (the uppermost of the Ordovician System). Episodes 29(3), 183-196.

Chen Xu, Rong Jia-yu, Mitchell, C.E., Harper, D.A.T., Fan Jun-xuan, Zhan Ren-bin, Zhang Yuan-dong, Li Rong-yu & Wang Yi 2000. Late Ordovician to earliest Silurian graptolite and brachiopod biozonation from the Yangtze region, South China, with a global correlation. Geological Magazine 137(6), 623-650.

Chlupáč, I. 1951a. Stratigrafické poměry králodvorských břidlic u Karlíka a Zadní Třebáně. Věstník Ústředního ústavu geologického 26, 194-212. [in Czech with Russian and English summary]

Chlupáč, I. 1951b. Profil králodvorskými břidlicemi (Ashgillian) u Velké Chuchle. Věstník Královské české společnosti nauk, Třída matematicko-přírodovědecká 1950(1), 1-10. [in Czech with English summary]

Chlupáč, I. 1953. Poznámky ke stratigrafii králodvorských břidlic v okolí Litně. Časopis Národního musea, Oddíl přírodovědný 122, 28-33. [in Czech]

Chlupáč, I., Havlíček, V., Kříž, J., Kukal, Z. & Štorch, P. 1992. Palaeozoikum Barrandienu (kambrium - devon). 292 pp. Vydavatelství Českého geologického ústavu, Praha. [in Czech]

Chlupáč, I., Havlíček, V., Kříž, J., Kukal, Z. & Štorch, P. 1998. Palaeozoic of the Barrandian (Cambrian to Devonian). 183 pp. Czech Geological Survey, Prague.

Cooper, R.A. & Fortey, R.A. 1982. The Ordovician Graptolites of Spitsbergen. Bulletin of the British Museum of Natural History, Geology Series 36(3), 157-302.

Cooper, R.A. & Sadler, P.M. 2012. The Ordovician Period. With a contribution from Hammer, O. & Gradstein, F.M., 489-523. In Gradstein, F.M., Ogg, J.G., Schmitz, M.D. & Ogg, G.M. (eds) The Geological Timescale 2012, Volume 1. 1176 pp. Elsevier B.V., Amsterdam.

Davies, K.A. 1929. Notes on the graptolite fauna of the Upper Ordovician and Lower Silurian. Geological Magazine 66, 1-27.View article

Elles, G.L. & Wood, E.M.R. 1906. A monograph of British graptolites. Part 5. Monograph of the Palaeontographical Society 60(288), lxxiii-xcvi, 181-216.

Elles, G.L. & Wood, E.M.R. 1907. A monograph of British graptolites. Part 6. Monograph of the Palaeontographical Society 61(297), xcvii-cxx, 217-272.

Elles, G.L. & Wood, E.M.R. 1908. A monograph of British graptolites. Part 7. Monograph of the Palaeontographical Society 62(305), cxxi-cxlviii, 273-358.

Fortey, R.A. & Cocks, L.R.M. 2005. Late Ordovician global warming - The Boda event. Geology 33(5), 405-408.View article

Fortey, R.A. & Cooper, R.A. 1986. A phylogenetic classification of the graptoloids. Palaeontology 29(4), 631-654.

Frech, F. 1897. Lethaea Geognostica. Lethaea Paleozoica 1, 11, Graptolithen. Schweizerbart, Stuttgart.

Goldman, D. & Bergström, S.M. 1997. Late Ordovician graptolites from the North American midcontinent. Palaeontology 40(4), 965-1010.

Goldman, D., Mitchell, C.E., Melchin, M.J., Fan Jun-xuan, Wu Shuang-ye & Sheets, D.H. 2011. Biogeography and mass extinction: extirpation and re-invasion of Normalograptus species (Graptolithina) in the Late Ordovician palaeotropics. Proceedings of the Yorkshire Geological Society 58, 227-246.

Gurley, R.R. 1896. North American graptolites: new species and vertical range. Journal of Geology 4, 63-102.View article

Gutiérrez-Marco, J.C., Robardet, M., Rábano, I., Sarmiento, G.N., San José Lancha, M.Á., Herranz Araújo, P. & Pieren Pidal, A.P. 2002. Ordovician, 31-49. In Gibbons, W. & Moreno, T. (eds) The Geology of Spain. 649 pp. Geological Society, London.

Hadding, A., 1915. Der Mittlere Dicellograptus-Schiefer auf Bornholm. Lunds Universitets Arsskrift, Afd. 2, 11(4), 1-40.

Hall, J. 1865. Graptolites of the Quebec group. Figures and descriptions of Canadian organic remains. Decade II. 151 pp. Geological Survey of Canada, Ottawa.

Havlíček, V. 1981. Development of a linear sedimentary depression exemplified by the Prague Basin (Ordovician-Middle Devonian; Barrandian area - central Bohemia). Sborník geologických věd, Geologie 35, 7-48.

Havlíček, V. 1982. Ordovician in Bohemia: development of the Prague Basin and its benthic communities. Sborník geologických věd, Geologie 37, 103-136.

Havlíček, V. 1989. Climatic changes and development of benthic communities through the Mediterranean Ordovician. Sborník geologických věd, Geologie 44, 79-116.

Havlíček, V. & Marek, L. 1973. Bohemian Ordovician and its international correlation. Časopis pro mineralogii a geologii 18(3), 225-232.

Havlíček, V. & Vaněk, J. 1966. The Biostratigraphy of the Ordovician of Bohemia. Sborník geologických věd, Řada P - Paleontologie 8, 7-67.

Hisinger, W. 1837. Lethaea Suecica seu Petrificata Sueciae, Supplementum 1. 124 pp. Norstedt, Stockholm.

Hopkinson, J. 1871. On Dicellograptus, a new species of Graptolites. Geological Magazine 8, 20-26.View article

Hopkinson, J. & Lapworth, J. 1875. Descriptions of the graptolites of the Arenig and Landeilo rocks of St. David’s. Quarterly Jounal of the Geological Society of London 31, 631-672.View article

Jackson, D.E. 1971. Development of Glyptograptus hudsoni sp. nov. from Southampton island, North-West Territories, Canada. Palaeontology 14(3), 478-486.

Kaljo, D., Männik, P., Martma, T. & Nolvak, J. 2012. More about the Ordovician-Silurian transition beds at Mirny Creek, Omulev Mountains, NE Russia: carbon isotopes and conodonts. Estonian Journal of Earth Sciences 61(4), 277-294.View article

Koren’, T.N., Oradovskaya, M.M., Pylma, L.J., Sobolevskaya, R.F. & Chugaeva, M.N. 1983. The Ordovician and Silurian Boundary in the Northeast of USSR. 205 pp. Nauka Publishers, Leningrad. [in Russian with English abstract]

LaPorte, D.F., Holmden, C., Patterson, W.P., Loxton, J.D., Melchin, M.J. & Mitchell, C.E. 2009. Local and global perspectives on carbon and nitrogen cycling during the Hirnantian glaciation. Palaeogeography, Palaeoclimatology, Palaeoecology 276, 182-195.View article

Lapworth, C. 1873. Notes on the British graptolites and their allies I. On an improved classification of the Rhabdophora. Geological Magazine 1(10), 500-504, 555-560.View article

Lapworth, C. 1876. The Silurian System in the south of Scotland, 1-28. In Armstrong, J., Young, J. & Robertson, D. (eds) Catalogue of Western Scottish Fossils. 164 pp. Blackie and Son, Glasgow.

Lapworth, C. 1877. On the graptolites of County Down. Proceedings of the Belfast Naturalists’ Field Club, 1876-1877, 125-148.

Lapworth, C. 1879. On the geological distribution of the Rhabdopleura. Annals and Magazine of Natural History, Series 5(3), 245-257, 449-455; (4) 333-341, 423-431.

Lapworth, C. 1880. The Silurian sequence of Rhayader. Quarterly Journal of the Geological Society of London 56, 67-137.View article

Legrand, P. 1977. Contribution a l’étude des graptolites du Llandoverien inferieur de l’Oued In Djerane (Tassili N’ajjer oriental, Sahara algérien). Bulletin de la Société d’Histoire Naturelle de l’Afrique du Nord 67, 141-196.

Legrand, P. 1987. Modo de desarrollo del suborden Diplograptina (Graptolithina) en el Ordovício superior y en el Silúrico. Implicaciones taxonómicas. Revista Espanola de Paleontologia 2, 59-64.

Legrand, P. 1993. Graptolites d’age ashgillien dans la region de Chirfa (Djado, République du Niger). Bulletin du Centres Recherches Exploration-Production Elf Aquitaine 17, 435-442.

Legrand, P. 2011. On the Katian/Hirnantian boundary. application on the north-African border of Gondwana, 287-294. In Gutiérrez-Marco, J.C., Rábano, I. & García-Bellido, D. (eds) Ordovician of the World. Cuadernos del Museo Geominero 14. 679 pp. Instituto Geológico y Minero de Espana, Madrid.

Li Ji-jin 1984. Graptolites from the Xinling Formation (Upper Ordovician) of South Anhui. Memoirs of Nanjing Institute of Geology and Palaeontology, Academia Sinica 20, 145-194.

Li Ji-jin & Cheng Han-jun 1988. Graptolites from Nancheng Formation at Liangshan, Hanzhong of Shaanxi. Acta Palaeontologica Sinica 27(2), 164-178.

Loydell, D.K. 2007. Graptolites from the Upper Ordovician and Lower Silurian of Jordan. Special Papers in Palaeontology 78, 1-66.

Maletz, J., Carlucci, J. & Mitchell, C.E. 2009. Graptoloid cladistics, taxonomy and phylogeny. Bulletin of Geosciences 84(1), 7-19.View article

Marr, J.E. 1880. On the Predevonian rocks of Bohemia. Quarterly Journal of the Geological Society 36, 591-619.View article

McCoy, F. 1850. On some new genera and species of Silurian Radiata in the collection of the University of Cambridge. Annals and Magazine of Natural History 6, 270-290.

Melchin, M.J., Mitchell, C.E., Holmden, C. & Štorch, P. 2013. Environmental changes in the Late Ordovician - early Silurian: Review and new insights from black shales and nitrogen isotopes. The Geological Society of America Bulletin 125(11/12), 1635-1670.View article

Melchin, M.J., Mitchell, C.E., Naczk-Cameron, A., Fan Jun-xuan & Loxton, J. 2011. Phylogeny and adaptive radiation of the Neograptina (Graptoloida) during the Hirnantian mass extinction and Silurian recovery. Proceedings of the Yorkshire Geological Society 58(4), 281-309.View article

Mergl, M. 2011a. Earliest occurrence of the Hirnantia Fauna in the Prague Basin (Czech Republic). Bulletin of Geosciences 86(1), 63-70.View article

Mergl, M. 2011b. Faunal turnover near the Katian/Hirnantian boundary in the Prague Basin (Czech Republic), 359-366. In Gutiérrez-Marco, J.C., Rábano, I. & García-Bellido, D. (eds) Ordovician of the World. Cuadernos del Museo Geominero 14. 679 pp. Instituto Geológico y Minero de Espana, Madrid.

Mikhaylova, N.F. 1973. Graptolity verkhnego ordovika i nizhnego silura Kazakhstana, 14- 19. In Ivanovskiy, A.B. (ed.) Novoye v paleontologii Sibiri i Sredney Azii. 140 pp. Nauka, Novosibirsk. [in Russian]

Mitchell, C.E. 1987. Evolution and phylogenetic classification of the Diplograptacea. Palaeontology 30, 353-405.

Mitchell, C.E. , Goldman, D., Klosterman, S.L., Maletz, J., Sheets, H.D. & Melchin, M.J. 2007. Phylogeny of the Ordovician Diplograptoidea, 332-339. In Li Jun, Fan Jun-xuan & Percival, I. (eds) Proceedings of the 10th International Symposium of the Ordovician System, Nanjing, China, June 2007. Acta Palaeontologica Polonica 46 (Supplement).

Mitchell, C.E., Maletz, J. & Goldman, D. 2009. What is Diplograptus? Bulletin of Geosciences 84(1), 27-34.View article

Mitchell, C.E., Štorch, P., Holmden, C., Melchin, M.J. & Gutiérrez-Marco, J.C. 2011. New stable isotope data and fossils from the Hirnantian Stage in Bohemia and Spain: implications for correlation and paleoclimate, 371-378. In Gutiérrez-Marco, J.C., Rábano, I. & García-Bellido, D. (eds) Ordovician of the World. Cuadernos del Museo Geominero 14. 679 pp. Instituto Geológico y Minero de Espana, Madrid.

Mu En-zhi 1949. Two growth patterns of Climacograptus. Geological Review 14(4-6),185-188 [in Chinese].

Mu En-zhi 1963. Research in graptolite faunas of Chilianshan. Scientia Sinica 12(3), 347-371.

Mu En-zhi, Li Ji-jin, Ge Mei-yu, Chen Xu, Lin Yao-kun & Ni Yu-nan 1993. Upper Ordovician graptolites of Central China Region. Palaeontologia Sinica, New Series B, 189(29), 1-393. [in Chinese with English summary]

Mu En-zhi, Li Ji-jin, Ge Mei-yu, Lin Yao-kun & Ni Yu-nan 2002. Graptolites of China. 1205 pp. Science Press, Beijing. [in Chinese]

Mu En-zhi & Lin Yao-kun 1984. Graptolites from the Ordovician-Silurian boundary sections of Yichang area, S. Hubei, 45-73. In Nanjing Institute of Geology and Palaeontology, Academia Sinica (ed.) Stratigraphy and palaeontology of systematic boundaries in China, Ordovician-Silurian boundary 1. 516 pp. Anhui Science and Technology Publishing House, Hefei.

Ni Yu-nan 1984. Upper Ordovician graptolites from Baoshan, western Yunnan. Acta Palaeontologica Sinica 22, 320-327. [in Chinese with English summary]

Nicholson, H.A. 1867. On some fossils from the Lower Silurian rocks of south of Scotland. Geological Magazine 4(33), 107-113.View article

Obut, A.M. 1955. Graptolity, 136-139. In Nikiforova, O.I. (ed.) Polevoy atlas ordovikskoy i silurskoy fauny Sibirskoy platformy. Gosgeoltechizdat, Moskva. [in Russian]

Obut, A.M. & Sennikov, N.V. 1985. Osobennosti llandoveriyskikh planktonovykh soobschestv Sibirskoy platformy, 51-60. In Betekhtina, O.A. & Zhuravleva, I.T. (eds) Sreda i zhizn’ v geologicheskom proshlom (Paleobasseyny i ich obivateli). Trudy Instituta geologii i geofyziki 628. 137 pp. Nauka, Novosibirsk. [in Russian]

Obut, A.M. & Sobolevskaya, R.F. 1974. Graptolites, 85-159. In Palaeozoic reference sections of the North-East USSR. Nauka, Magadan.

Page, A.A., Zalasiewicz, J.A., Williams, M. & Popov, L.E. 2007. Were transgressive black shales a negative feedback modulating glacioeustasy in the Early Palaeozoic Icehouse?, 123-156. In Williams, M., Haywood, A.M., Gregory, F.J. & Schmidt, D.N. (eds) Deep-Time Perspectives on Climate Change: Marrying the Signal from Computer Models and Biological Proxies. The Micropalaeontological Society, Special Publications. 589 pp. The Geological Society, London.

Perner, J. 1895. Études sur les Graptolites de Boheme, IIieme Partie. Monographie des Graptolites de l’Étage D. 31 pp. Raimond Gerhard, Leipsic, Prague.

Přibyl, A. 1949. Revise českých ordovických diplograptidů a glossograptidů. Rozpravy České akademie věd a umění, Třída II 59, 1, 1-48. [in Czech]

Rickards, R.B. 2002. The graptolitic age of the type Ashgill Series (Ordovician), Cumbria, UK. Proceedings of the Yorkshire Geological Society 54(1), 1-16.View article

Ross, R.B. & Berry, W.B.N. 1963. Ordovician graptolites of the Basin Ranges in California, Nevada, Utah and Idaho. U.S. Geological Survey Bulletin 1134, 1-177.

Ruedemann, R. 1908. Graptolites of New York, Part 2. Graptolites of higher beds. Memoirs of the New York State Museum of Natural History, Albany 11, 457-583.

Ruedemann, R. 1925. The Utica and Lorraine Formations of New York, Pt. II: Systematic Paleontology 1 (Plants, Sponges, Corals, Graptolites, Crionoids, Worms, Bryozoans, Brachiopods). New York State Museum Bulletin 262, 1-171.

Shaw, F.C. 2000. Trilobites of the Králův Dvůr Formation (Ordovician) of the Prague Basin, Czech Republic. Bulletin of the Czech Geological Survey 75(4), 371-404.

Skoglund, R. 1963. Uppermost Viruan and Lower Harjuan (Ordovician) stratigraphy of Västergötland and Lower Harjuan graptolite fauna of central Sweden. Bulletin of Geological Institutions of the University Uppsala 42, 1-55.

Sobolevskaya, R.F. 1974. New Ashgill graptolites in the middle flow basin of the Kolyma River, 63-71. In Obut, A.M. (ed.) Graptolites of the USSR. 160 pp. Nauka, Novosibirsk. [in Russian]

Sobolevskaya, R.F. 2011. Atlas of the Paleozoic fauna of Taimyr. Part II. The Ordovician and Silurian graptolites. Trudy NIIGA-VNIIOkeanologia 221. 281 pp. FGUP VIINOkeanologia im. I.S. Gramberga, Sankt-Peterburg. [in Russian]

Stewart, S. & Mitchell, C.E. 1997. Anticostia, a distinctive new Late Ordovician “glyptograptid” (Diplograptacea, Graptoloidea) based on three-dimensionally preserved specimens from Anticosti Island, Quebec. Canadian Journal of Earth Sciences 34, 215-228.View article

Štorch, P. 1989. Late Ordovician graptolites from the upper part of the Králův Dvůr Formation of the Prague Basin (Barrandian, Bohemia). Věstník Ústředního ústavu geologického 64(3), 173-186.

Štorch, P. & Mergl, M. 1989. Králodvor/Kosov boundary and the late Ordovician environmental changes in the Prague Basin (Barrandian area, Bohemia). Sborník geologických věd, Geologie 44, 117-153.

Štorch, P., Mitchell, C.E., Finney S.C. & Melchin, M.J. 2011. Uppermost Ordovician (upper Katian-Hirnantian) graptolites of north-central Nevada, U.S.A. Bulletin of Geosciences 86(2), 301-386.View article

Štorch, P. & Serpagli, E. 1993. Lower Silurian Graptolites from Southwestern Sardinia. Bolletino della Societa Paleontologica Italiana 32, 3-57.

Vandenbroucke, T., Rickards, R.B. & Verniers, J. 2005. Upper Ordovician chitinozoan biostratigraphy from the type Ashgill area (Cautley district) and the Push Gill section (Dufton district, Cross Fell Inlier), Cumbria, Northern England. Geological Magazine 142(6), 783-807.View article

Wang Ju-de 1974. Graptolites, 731-736. In Yunnan Bureau of Geology (ed.) Fossil Atlas of Yunnan, Part 1. Peoples Publishing House of Yunnan, Kunming. [in Chinese]

Wang Xiao-feng 1978. Ordovician Graptolites, 192-210. In Stratigraphy Research Group of the Yangtze Gorges, Hubei Bureau of Geology (ed.) Sinian to Permian stratigraphy and palaeontology of eastern Yangtze Gorges. 381 pp. Geological Publishing House, Beijing. [in Chinese]

Wang Xiao-feng, Zeng Qing-luan, Zhou Tian-mei, Ni Shi-zhao, Xu Guang-hong, Sun Quan-ying, Li Zhi-hong, Xiang Li-wen & Lai Cai-gen 1983. Latest Ordovician and earliest Silurian faunas from the eastern Yangtze Gorges, China with comments on Ordovician-Silurian boundary. Bulletin of Yichang Institute of Geology and Mineral Resources, Chinese Academy of Geological Sciences 6, 126-196.

Whittington, H.B. 1955. Additional new Ordovician graptolites and a chitinozoan from Oklahoma. Journal of Paleontology 29, 837-851.

Williams, S.H. 1981. Form and mode of life of Dicellograptus (Graptolithina). Geological Magazine 118, 401-408.View article

Williams, S.H. 1982. The late Ordovician graptolite fauna of the Anceps Bands at Dob’s Linn, southern Scotland. Geologica et Palaeontologica 16, 29-56.

Zalasiewicz, J.A., Taylor, L., Rushton, W.A., Loydell, D.K., Rickards, R.B. & Williams, M. 2009. Graptolites in British stratigraphy. Geological Magazine 146, 785-850.View article